3. Benstonea alticola (Holttum & H. St. John) Callm. & Buerki, comb. nova. ≡ Pandanus alticola Holttum & H. St. John in Pacific Sci. 16: 218. 1962. Typus: MALAYSIA. Johore (Peninsular Malaysia): Sungei Kayu, Mawai-Jemaluang rd., II.1935, Corner s.n. (holo-: SING [SING0169801]!). Distribution and ecology. – Benstonea alticola is found in montane forest and is an epiphyte growing to 12 m above ground, or a terrestrial shrub on limestone domes in Peninsular Malaysia (Bukit Takun, Selangor; Gua Musang, Kelantan) and Borneo (Sarawak) at an altitude of c. 700-1500 m. Observations. – This facultative epiphytic shrub, with one large clump of leaves on a very short stem, has only a single terminal syncarp. Abroader species concept, including other facultative epiphytes recognized here as distinct species such as B. inquilina (B. C. Stone) Callm. & Buerki, may ultimately prove to be more appropriate, but further fieldwork will be required to determine the preferable circumscription of species.

Published as part of Callmander, Martin W., Lowry Ii, Porter P., Forest, Félix, Devey, Dion S., Beentje, Henk & Buerki, Sven, 2012, Benstonea Callm. & Buerki (Pandanaceae): characterization, circumscription, and distribution of a new genus of screw-pines, with a synopsis of accepted species, pp. 323-345 in Candollea 67 (2) on page 330, DOI: 10.15553/c2012v672a12, http://zenodo.org/record/5790002