Iodes scandens (Becc.) Utteridge & Byng Fig. 15.10–15.17 Material examined Specimen used for endocarp and fruit description NEW GUINEA • s. loc; s.d.; s. col. s.n.; MO. Description FRUIT. Elliptical; epicarp puberulent. Calyx persistence and mesocarp thickness unknown. Length 20 mm, width 15 mm, thickness 15 mm. ENDOCARP. Brown, elliptical in lateral view, lenticular in transverse section, length ca 14.7 mm, width ca 12.9 mm, thickness ca 11.2 mm. Keel surrounding the endocarp in the plane of symmetry, with the thicker margin containing the primary vascular bundle embedded within the endocarp wall. Apex rounded and asymmetrical; base rounded, slightly asymmetrical. Outer surface of the endocarp ridged with two main longitudinal ridges; the median ridge runs from the point of the base up the half of the endocarp length, then divides into several smaller ridges and a lateral ridge, which runs from the subbase up to the subapex. Ridges rectangular, thin, with a median channel, delimiting a reticulate pattern enclosing 13–18 irregular ridges with arbuscular freely ending ridgelets. Vasculature of the endocarp surface in the channels of the ridges. Endocarp possessing a symmetrical pair of horn-like protrusions compressed on the keel, positioned eccentrically and subapically on the endocarp faces, each with a central pit. Endocarp wall 502–564 µm thick excluding ridges (652–672 µm including ridges). Endocarp wall with three cell layers: outermost layer with 11–13 rows of anticlinally oriented cells, cells 14.0– 43.1 µm in length, followed by a layer with 6–8 rows of periclinally oriented cells, cells 21.1–25.6 µm in width; innermost layer with one row of periclinally oriented cells, cells 9.3–9.8 µm in width, covering the locule surface, apparently with rounded papillae (detail of the surface poorly preserved). Locule lacunate.

Published as part of Rio, Cédric Del, Stull, Gregory W. & Franceschi, Dario De, 2020, Survey of the fruits and endocarps of Icacinaceae (Lamiids, Icacinales), pp. 1-130 in European Journal of Taxonomy 645 on page 44, DOI: 10.5852/ejt.2020.645, http://zenodo.org/record/3829651