Although the ceremonial center of Vilcanota was called the third most important temple in the Inka empire in the sixteenth century, its exact location and meaning have remained matters of conjecture. In this article I examine historical and archaeological information which demonstrates that the temple was located at the pass of La Raya. Ecological and ethnographic data from the region support the conclusion that the temple was built at La Raya because of the area's association with sacred rivers and mountains which were in turn linked with fertility concepts, the birth of the sun, and an ecological/political boundary. Together these factors made the place of special significance in Inka religion.