Information on the aerial distribution of Tipulinae is fragmentary and scattered, the only detailed studies being the light-trap investigations of nocturnal activity by Pinchin & Anderson (1936) and Robertson (1939) and the recent suction-trap investigation of woodland species by Service (1973). In the present investigation, suction traps were used to trap Tipulinae at five localities in Scotland. Data on the seasonal occurrence and on the relative numbers of the species caught at four of these sites have been published already (Dobson 1973, 1974); here the vertical distribution of these species, their sex-ratios and the reproductive condition of the females will be considered.