With the help of relatively few conservative crop parameters, AquaCrop simulates final crop yield in four steps that are easy to understand, which makes the modeling approach transparent. The steps consist in the simulation of development of the green crop canopy cover, crop transpiration, above-ground biomass, and final crop yield. Temperature and water stresses directly affect one or more of the above processes. Nutrient deficiencies and salinity effects are simulated indirectly by moderating canopy cover development over the season, and by reducing crop transpiration and the normalized water productivity. The effect of CO2 concentration on biomass is simulated by altering the normalized water productivity. The model requires a relatively small number of explicit parameter values and mostly intuitive input variables. The paper describes the essence of AquaCrop Version 4.0, applications and parameterization of crops, crop responses to elevated CO2 concentration, soil fertility and salinity, and further model developments.