Groutiella tuberculata B.H. Allen & I.H. Holz (Fig. 12) Monographs in Systematic Botany from the Missouri Botanical Garden 90: 555 (Allen 2002). SPECIMENS EXAMINED. — Costa Rica. San José, Cordillera de Talamanca, 4 km N San Gerardo de Dota and 1 km W of Jaboncillo, Holz CR 99-981 (iso-, MO). DISTRIBUTION AND HABITAT. — Central America. On bark of branches, 2900 m. DESCRIPTION For description, see Allen (2002). COMMENTS Groutiella tuberculata is easy to be recognized for its consistently papillose basal leaf cells. Groutiella tomentosa is the other species of the genus with papillose basal leaf cells. The differences between them provided in the discussion of G. tomentosa. G. husnotii and G. macrorrhyncha are similar to G. tuberculata in having lanceolate and cripate leaves, however, they have smooth basal leaf cells, while, G. husnotii differs in a longer limbidia extending to 2/3 or more of leaf length, and G. macrorrhyncha differs in stoutly excurrent costa.

Published as part of Yu, Ning-Ning & Jia, Yu, 2023, A world revisionary study of the genus Groutiella Steere (Orthotrichaceae, Bryopsida), pp. 161-182 in Cryptogamie, Bryologie 20 (7) on page 178, DOI: 10.5252/cryptogamie-bryologie2023v44a7, http://zenodo.org/record/8155687