Clusia flava (Meigen, 1830) 38 specimens: 15 males (m), 19 females (f), 4 empty puparia; host trees: Fagus (26 records), Betula (7), Alnus (1), Fraxinus (1), Picea (1), Pinus (2). Scotland: 2m, 2f, Midlothian, Arniston, 26.v.1995, ex Fagus log, DH; 1f, 2 empty puparia, Midlothian, Newbattle Abbey, 22–27.v.1995, ex fallen Fagus branch, GER; 1f, Perthshire, Balnagard, 20.iii.1997, ex fallen Alnus, GER; 2m, 2f, Perthshire, Fungarth Wood, 26.xi.1997, ex fallen Betula, GER; 1m, Perthshire, Fungarth Wood, 1.iv.1998, ex fallen Betula, GER; 1m, 1f, Perthshire, Den of Alyth, no date, ex Fagus stump, IM; 1f, Morayshire, nr Ordiquish, 10.v.1999, ex Picea log, IM; 1f, Ross and Cromarty, Rassal Ashwood NNR, 26.v.2008, ex Fraxinus, DH; 2 empty puparia, Inverness-shire, Grantown-upon-Spey, Cemetery Wood, 12.viii.2012, ex Pinus stump, GW. England: 1m, 1f, Buckinghamshire, Burnham Beeches, 20.iv.1998, ex fallen Betula, GER; 3m, 6f, Buckinghamshire, Burnham Beeches, 21.iv.1998, ex Fagus log, GER; 5m, 2f, Buckinghamshire, Ashridge, 22.iv.1998, ex Fagus log, GER; Norway: 1f, Brana Kolhere, nr Oslo, 19.v.2005, ex fallen Fagus, GER. Previous host tree records: Fagus, Betula (Smith 1950, 1989). Clusia tigrina (Fallén 1820) 12 specimens: 2 males, 3 females, 7 empty puparia, all from Fagus. England: 7 empty puparia, Berkshire, Windsor Forest, 15.vii.1999, ex fallen Fagus, PJC; France: 2m, 3f, Pyrenees-Orientales, Reserve Naturelle de la Massane, near Banyuls-sur-mer, 8.v.2003, ex fallen Fagus, GER. Previous records: Fagus (Krivoscheina and Mamaev 1967, Chandler 1973); Clusiodes albimanus (Meigen, 1830) 69 specimens: 31 males, 38 females; host trees: Betula (37), Fagus (13), Fraxinus (13), Alnus (2), Populus (3), Salix (1). Scotland: 1f, Midlothian, Crichton Glen, 21.iii.1994, ex Fraxinus log, GER; 1m, 1f, Perthshire, Methan Woods,?. vi.1994, ex Fagus stump, IM; 3f, Clackmannanshire, nr Alva, Myretoun Wood, 26.ii.1995, ex fallen Fraxinus, GER; 2m, 2f, Midlothian, Edinburgh, Hermitage of Braid, 7.v.1995, ex fallen Fagus, GER; 1m, Midlothian, Arniston, 26.v.1995, ex fallen Fagus, IM; 1m, 2f, Inverness-shire, Kincraig, Speybank, 7.v.1996, ex fallen Betula, GER; 1m, Ross and Cromarty, Glen Cannich, 18.v.1996, ex Betula log, DH; 1f, Inverness-shire, Kincraig, Speybank, 19.v.1996, ex fallen Betula, GER; 1f, Perthshire, Balnagard, 20.iii.1997, ex fallen Alnus, GER; 1f, Ross and Cromarty, Braelangwel Wood, 18.v.1997, ex fallen Betula, GER; 4m, Ross and Cromarty, Rassal Ashwood NNR, 24.iv.1997, ex fallen Fraxinus, GER; 1f, Sutherland, The Mound, Loch Fleet, 25.iv.1997, ex Alnus branch, GER; 1m, 1f, Ross and Cromarty, Loch Achility, 27.vi.1998, ex Populus tremula L. branch, DH; 1m, Midlothian, Edinburgh, Hermitage of Braid, 25.v.1999, ex Fraxinus log, DH; 1f, Midlothian, Edinburgh, Hermitage of Braid, 25.v.1999, ex Fagus log, DH; 1f, Aberdeenshire, Linn of Dee, 31.v.1999, ex fallen Betula, GER. 2m, 4f, Ross and Cromarty, Rassal Ashwood NNR, 26.v.2008, ex Fraxinus branches, DH; 1m, Ross and Cromarty, 28.v.2008, ex Betula branch, DH; 1m, Perthshire, Dunkeld, Fungarth, 14.x.2007, ex Betula branch, DH. England: 1m, 7f, Buckinghamshire, Burnham Beeches, 30.iv.1997, ex Betula log, GER; 3m, 2f, Buckinghamshire, Burnham Beeches, 21.iv.1998, ex Betula log, GER; 2f, Buckinghamshire, Burnham Beeches, 20–21.iv.1998, ex fallen Fagus log, GER 3m, 3f, Buckinghamshire, Burnham Beeches, 23.iv.1998, ex Betula log, GER; 2m, 1f, Buckinghamshire, Ashridge, 22.iv.1998, ex Fagus log, GER; 1m, Buckinghamshire, Burnham Beeches, 23.iv.1998, ex fallen Salix, GER; Finland: Kainuu, Kuhmo: 1f, Jauhovaara, 17.v.2004, ex Betula, DH; 1m, Urpovaara NR, 30.v.2007, ex Betula branch, DH; 1m, Ulvinsalon NR, 31.v.2007, ex Betula branch, DH; 1f, Honkavaara, 1.vi.2007, ex Betula branch, DH; 2m, Puolanka, Baljakka NR, 4.vi.2007, ex Betula branch, DH; 1m, Valkeavaara, 3.vi.2007, ex Betula branch, DH. Previous records: Betula (Richards 1934); Fraxinus (Withers 1985). Clusiodes apicalis (Zetterstedt, 1848) 47 specimens: 19 males, 28 females, host trees: Betula (42), Alnus (4), Populus (1). Scotland: 1f, Inverness-shire, Kincraig, Speybank, 7.v.1996, ex fallen Populus, GER; 1m, Inverness-shire, Kincraig, Speybank, 19.v.1996, ex Betula branch, DH; 2m, 5f, Inverness-shire, Kincraig, Speybank, 19.v.1996, ex fallen Betula, GER; 1f, Inverness-shire, Kincraig, Speybank, 19.v.1996, ex fallen Betula, AEW; 6m, 6f, Perthshire, nr Balnagard, 20.iii.1997, ex Betula branch, GER; 2m, 2f, Sutherland, The Mound, Loch Fleet, 25.vi.1997, ex fallen Alnus, GER; 5f, Ross and Cromarty, Braelangwel Wood, 18.v.1997, ex Betula branch, DH; 1m, Perthshire, Fungarth, 1.iv.1998, ex fallen Betula, GER; 1m, 2f, Perthshire, Camghouran, Coille Mhor, 10.v.1998, ex Betula branch, DH; 1m, Perthshire, nr Tummel Bridge, 10.v.1998, ex fallen Betula, GER. Finland: Kainuu, Kuhmo: 1m, 3f, Jauhovaara, 64' 03"N, 29'10"E, 17.v.2004, ex fallen Betula, GER; 1f, Elimyssalo, 64' 15"N, 30'33"E, 23.v.2004, ex fallen Betula, GER; 1m, Ulvinsalon NR, 63'97"N, 30'32"E, 31.v.2007, ex Betula branch, DH; 2m, Honkavaara, 63'89"N, 29'68"E, 1.vi.2007, ex Betula branch. Russia: Karelia, Kostomuksha: 1m, 1f, west of Kostomukska, 64'59"N, 30'32"E, 18.v.2004, ex Betula branch, DH; 1m, Luvajarvi, 64'49"N, 30'70"E, 18.v.2004, ex Betula branch, DH. Previous records: Betula (Teskey 1976). Clusiodes caledonicus (Collin, 1912) 13 specimens: 8 males, 5 females all from Pinus. Scotland: 1m, 2f, Inverness-shire, Abernethy, Bognachruie, 3.v.1994, ex wet detritus under bark of dead, standing Pinus, GER; 1f, Ross and Cromarty, Shieldag, no date, ex fallen Pinus, IM; 4m, 2f, Inverness-shire, Abernethy, nr Forest Lodge, 12.vi.1997, ex Pinus stump, GER; 3m, Inverness-shire, Abernethy, nr Forest Lodge, 12.vi.1997, ex Pinus stump, DH. Previous records: Populus tremula L. (Tuomikoski 1933) Clusiodes freyi Tuomikoski, 1933 20 specimens, 3 male, 17 females, Betula (19), Picea (1). Finland: Kainuu, Kuhmo: 1f, Jauhovaara, 64'03"N, 29'10"E, 17.v.2004, ex Picea branch, DH; 1m, 8f, Elimyssalo, 64' 15"N, 30'33"E, 23.v.2004, ex fallen Betula, GER; 1f, Elimyssalo, 2.vi.2007, ex Betula branch, DH; 1f, Ulvinsalon NR, 63'97"N, 30'32"E, 21.v.2004, ex Betula branch, DH; 1m, 1f, Issakka, 63'88"N, 30'11"E, 21.v.2004, ex Picea, DH; 2m, 3f, Honkavaara, 63'89"N, 29'68"E, 1.vi.2007, ex Betula, DH. Russia: Karelia, Kostomuksha: 3f, west of Kostomuksha, 64'59"N, 30'32"E, 18.v.2004, ex Betula, DH; 1f, Luvajarvi, 64'49"N, 30'70"E, 18.v.2004, ex Betula, DH. Previous records: apparently not reared. Clusiodes geomyzinus (Fallén, 1823) 14 specimens: 9 males, 5 females, host trees: Picea (11), Pinus (2),? Betula (1). Finland: 7m, 4f, Kainuu, Kuhmo, Ulvinsalon Rasivaara, 63' 97"N, 30'32"E, 21.v.2004, ex Picea snag, GER; Russia: Karelia, Kostomuksha: 1f, Lake Korpangi, 64'75"N, 30'76"E, 18.v.2004, ex? Betula, SH; 2m, Luvajarvi, 64'49"N, 30'70"E, 18.v.2004, ex Pinus branch, DH. Previous records: apparently not reared, but adults associated with Pinus (Tuomikoski 1933, Stubbs 1982), Betula and P. t re m u l a (Tuomikoski 1933). Clusiodes gentilis (Collin, 1912) 49 specimens: 24 males, 25 females, host trees: Betula (38), Pinus (6), Fagus (2), Salix (1), Ulmus (1). Scotland: 3m, 3f, Perthshire, Fungarth, 26.ii.1997, ex fallen Pinus, GER;1f, Midlothian, Roslin Glen, 8.iii.1997, ex Fagus branch, DH; 1m, Inverness-shire, Grantown, Kylintra wood, 4.vi.2001, ex Pinus branch, DH; 1m, East Lothian, Pease Bay, 27.i.2008, ex Ulmus branch, DH. England: 19m, 18f, Buckinghamshire, Burnham Beeches, 19–22.vi.1998, ex Betula, GER; 2f, Buckinghamshire, Burnham Beeches, 21.vi.1998, ex Fagus branch, GER; 1f, Buckinghamshire, Burnham Beeches, 21.vi.1998, ex Salix branch, GER. Previous records: Betula, Salix (Withers 1985). Clusiodes microcerus Stackelberg, 1955 9 specimens: 3 males, 6 females, all from Betula. Finland: Kainuu, Kuhmo: 3m (reared, but no puparia), 4f (with puparia), Jauhovaara, 64' 03"N, 29'10"E, 17.v.2004, ex Betula branch, DH; 1f, Ulvinsalon NR, 21.v.2007, ex Betula branch, DH; 1f, Honkavaara, 1.vi.2007, ex Betula branch, DH. Previous records: apparently not reared. Clusiodes ruficollis (Meigen, 1830) 18 specimens: 9 males, 9 females, host trees: Fagus (5), Betula (3), Fraxinus (2), unknown (8). Scotland: 5m, 3f, Fife, St. Andrews, 12.x.1945, ex dead wood, tree species not recorded, DJ; 2f, Midlothian, Crichton Glen, 21.iii.1994, ex Fraxinus, DH; 1m, 3f, Midlothian, Edinburgh, Hermitage of Braid, 7.v.1995, ex fallen Fagus, GER; 1m, Inverness-shire, Aviemore, Craigellachie, 12.iv.1997, ex Betula branch, DH; 1m, Midlothian, Edinburgh, Hermitage of Braid, 4.v.1998, ex Fagus branch, DH; 1f, Perthshire, Struan, 9.v.1998, ex fallen Betula, GER; England: 1m, Buckinghamshire, Burnham Beeches, 21.iv.1998, ex Betula branch, GER. Previous records: Alnus, Betula (Tuomikoski 1933); P. t re m u l a (Greve and Midtgaard 1986); Carpinus (Withers 1985); Betula, Fagus (Rohácek 1995) Clusiodes verticalis (Collin, 1912) 7 specimens: 3 males, 4 females, host trees: Betula (2), Alnus (1), Fagus (1), Fraxinus (1), Pinus (1), Salix (1). Scotland: 1m, Midlothian, Crichton Glen, vi.1995, ex Fraxinus branch, GER; 1f, Inverness-shire, Glen Cannich, 18.v.1996, ex Betula branch, DH; 1m, Midlothian, Hermitage of Braid, 9.vi.1996, ex Pinus branch, DH; 1f, Perthshire, Balnagard, 20.iii.1997, ex fallen Alnus, GER; England: 1m, Buckinghamshire, Burnham Beeches, 21.vi.1998, ex Betula branch, GER; 1f, Buckinghamshire, Burnham Beeches, 21.vi.1998, ex Fagus branch, GER; 1f, Buckinghamshire, Burnham Beeches, 23.vi.1998, ex Salix branch, GER. Previous records: 'old grass heap' (Smith 1989); Betula (Withers 1985).

Published as part of Horsfield, David, 2013, Development sites and early stages of eleven species of Clusiidae (Diptera) occurring in Europe, pp. 401-427 in Zootaxa 3619 (4) on pages 403-405, DOI: 10.11646/zootaxa.3619.4.1, http://zenodo.org/record/223688