2.3 Colocasiomyia sabahana species complex, new Diagnosis. Cibarial, posterior sensilla absent (ch.6-1; Fig. 4C). Phallal sheath gently curved dorsad and slightly projected triangularly at dorsobasal corner in lateral view (ch.40-2, 41-1; Figs 11F, 24I). Shared characters. Supracervical setae approximately 4 per side, distally more or less curved, longer than inner occipital setae. Anterior reclinate orbital seta small, situated behind proclinate orbital seta; additional, interfrontal setulae on fronto-orbital plate present (ch.2-0). Eye with stout, dense interfacetal setulae (ch.3-1). Distance between antennal sockets wider than half of socket width (ch.4-0). Cibarial, medial sensilla 3–4 per side, in parallel rows narrower than sensilla campaniformia (ch.5-2; Fig. 4C). Anterior dorsocentral setae behind transverse suture (ch.7- 0). Prescutellar acrostichal setae absent (ch.8-1); acrostichal setulae in 4 rows (ch.9-0). Katepisternal setae small, shorter than postpronotal setae (ch.10-1). Apical scutellar setae nearer to each other than to basal scutellar seta. Costal setae in middle row all weak, trichoid (ch.12-2). Patch covered with only minute pubescence absent on anterodorsal portion of fore tarsomere I (ch.13-1). Male abdominal sternites III–V wider than long (ch.16,17,18-0); V medially concaved on posterior margin. Female abdominal sternite III as long as wide (ch.19-0); V longer than wide (ch.21-1). Male abdominal sternite VI not pubescent at least partly, with a pair of divergent processes longer than 0.1 mm, widely separated from each other, bearing prominent seta on apex and small setae on submedial to distal portion (ch.23,25-0, 24,26,27-1; Fig. 9L,M). Epandrium pubescent except for anterior to ventral margin and anteroventral elongation; setae on posteroventral portion as thick as cercal setae (ch.32-0); phragma small (narrower than epandrial anteroventral corner) expansion on antero-subapical margin (ch.33-2). Surstylus vestigial, very narrow process, apically with 1 or 2 tiny, trichoid seta(e) (ch.34,35-1). Cercus pubescent only on dorsal half; ventral/apical portion more or less differentiated from upper portion, with many small setae (ch.38-0). Phallal sheath not pubescent, as thick as wide, apically not bilobed (ch.42-0, 43,44-1; Figs 11F,G, 24I,J); phallapodeme sclerotized plate directed nearly along or slightly oblique to phallal axis (ch.45-1, 46-0; Figs 11F, 24I). Gonocoxites long, narrow slips (Figs 11F, 24I). Epiproct and hypoproct with pubescence (ch.48-0). Oviscapt shorter than phallus (apodeme + sheath), with ovisensilla only on distal portion occupying less than 1/2 of total length (ch.49-0, 51-1). Puparium (3rd instar larva): segments with stout spicules on ventral surface; anterior spiracle with stalk ending in a whorl of 19–25 long branches; stalk as long as posterior spiracle and longer than longest branch; caudal segments not so elongate, with many small spicules, ending in a V-shaped pair of long posterior spiracles (Fig. 24M,N). Mouth hook moderately, triangularly expanded medioventrally in lateral view; distal blade as long as basal portion, apically pointed, gently curved downward, with two rows of small, acute teeth on submedial to subapical portion of ventral margin (Fig. 24O,P). Included species. Colocasiomyia sabahana sp. nov. and C. sarawakana sp. nov.

Published as part of Takano, Kohei Takenaka, Gao, Jian-Jun, Hu, Yao-Guang, Li, Nan-Nan, Yafuso, Masako, Suwito, Awit, Repin, Rimi, Pungga, Runi Anak Sylvester, Meleng, Paulus Ak, Kaliang, Clement Het, Chong, Lucy & Toda, Masanori J., 2021, Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species, pp. 1-70 in Zootaxa 5079 (1) on page 37, DOI: 10.11646/zootaxa.5079.1.1, http://zenodo.org/record/5766402