Leptolalax sabahmontanus sp. nov. (Figures 2, 3, 4) Holotype. BORNEENSIS 22684, an adult male from Silau Silau trail along Sungai Silau-Silau (06o00'N, 116o32'E, alt. 1450 m a.s.l.), Kinabalu National Park (KNP), Ranau District, Sabah, Malaysia, collected on 31 July 2005 by M. Matsui. Paratypes. BORNEENSIS 22683, 22685, 22687–22691, 22701, KUHE 48014–48016 (former BORNEENSIS 22681, 22682, 22686), 11 males data, same as the holotype. BORNEENSIS 22757, 22760, KUHE 48017 (former BORNEENSIS 22756), three males from the type locality, collected on 2 August 2005 by M. Matsui, P. Yambun, K. Nishikawa, and T. Shimada. SP 26736, 26737, two males from the type locality, collected on 14 August 2008 by Majin Silim. BORNEENSIS 22748, 22749, two males from Pandanus trail (06o00'N, 116o32'E, 1500 m a.s.l.), KNP, collected on 1 August 2005 by M. Matsui, P. Yambun, K. Nishikawa, and T. Shimada. KUHE 48018 (former BORNEENSIS 22784), BORNEENSIS 22785, 22786, three males from junction of Sg. Liwagu and Sg. Silau-Silau (06o00’ N, 116o33’ E, 1400 m a.s.l.), KNP, collected on 5 August 2005 by M. Matsui, P. Yambun, K. Nishikawa, and T. Shimada. SP 26740, an adult female from Bundu Tuhan view trail, KNP, collected on 14 August 2008 by S. David. BORNEENSIS 22980, 23029, 23034, three males from Sayap (06o11’ N, 116o34’ E, 950m a.s.l.), KNP, Kota Belud District, collected on 12 August 2005 by M. Matsui, Y. B. Paul, K. Nishikawa, and T. Shimada. Referred specimens. BORNEENSIS 8711, 8726, 8729, three males from Trail 4 (05o29’ N, 116 o01’ E, 754 m a.s.l.) of UMS 2002 expedition to the Crocker Range, Ulu Kimanis, Crocker Range National Park (CRNP), Tambunan District, collected on 9 September 2002 by M. Matsui. BORNEENSIS 8179, one male from Trail 5 (05o29’ N, 116o01’ E, 820 m a.s.l.) of Ulu Kimanis, CRNP, collected on 1 September 2002 by M. Matsui. BORNEENSIS 8673, one male collected on 7 September 2002 by M. Matsui, and KUHE 48020 (former BORNEENSIS 23470), one male collected on 19 August 2005 by M. Matsui, from Trail 11 (05o27’ N, 116o03’ E, 1427 m a.s.l.) of Ulu Kimanis, CRNP. KUHE 48019 (former BORNEENSIS 8855), one male collected on 12 December 2003, BORNEENSIS 12454, one male collected on 5 August 2003 by M. Matsui, BORNEENSIS 12632–12634, three males collected on 9 August 2003 by M. Matsui, K. Nishikawa, and T. Shimada, and BORNEENSIS 23482, one male collected on 20 August 2005 by M. Matsui, all from Mahua, CRNP (05o48’ N, 116o24’ E, 1063 m a.s.l.). Etymology. The specific epithet is from Sabah and a Latin adjective montanus denoting mountain, alluding to the fact that the species inhabits on mountains of Sabah, northern Borneo. Diagnosis. The new species is assigned to Leptolalax on the basis of the following: lack of vomerine teeth; fingers rounded in tips and lacking webbing; inner palmar tubercle elevated, not extending to the first finger; subarticular tubercles indistinct, replaced by low callous tissue; lack of nuptial pad; toe webbing basal; lack of outer metatarsal tubercle; snout tip with white vertical bar (Dehling & Matsui 2013). A medium-sized form of the genus, adult males about 25–28 mm in SVL; snout rounded in both ventral view and lateral view; interorbital space narrower than upper eyelid; male vocal sac not bipartite; toe webbing basal; dorsal skin of trunk and head smooth with tiny tubercles; pectoral gland small and indistinct; supraaxillary gland and ventrolateral glandular ridges absent; venter spotted; advertisement call consisting of long series of 1–149 notes, each composed of one to two indistinct pulse groups, dominant frequency at 6.75–7.35 kHz, without prominent frequency modulation. Most similar to L. dringi, but differing in smaller body size, relatively smaller eye-tympanum distance and inner metatarsal tubercle, and larger interorbital distance, upper eyelid width, snout length, eye length, and forearm and hand length compared to L. dringi. Description of holotype (measurement in mm). SVL 26.3; head and body moderately stocky (Figures 3); head longer (10.7) than broad (9.7) and wider than trunk; snout rounded in both ventral view and lateral view, slightly protruding beyond lower jaw; eye large (4.1) subequal to snout length (4.0); pupil vertical; canthus rostralis distinct, slightly concave in dorsal view between eye and nostril; convex between nostril and tip of snout; loreal region oblique, concave; nostril lateral, below canthus, distinctly closer to eye (2.1) than to tip of snout; internarial space (2.5) narrower than interorbital distance (3.0), latter slightly narrower than upper eyelid (3.2); pineal spot invisible; tympanum distinct, rounded, diameter (2.4) three-fifths that of eye, and separated from eye by about onethird tympanic diameter (0.9); symphysial knob on anteriormost part of mandible; vomerine ridge and teeth absent; tongue large, broad, notched distally, without papillae; a median, subgular vocal sac in the mid-ventral line; vocal openings small, just posterior to rictus, close to eustachian tube openings. Forelimb slender, moderately long (19.7); hand and forearm long (15.0); fingers long and slender, without webbing or lateral fringe of skin (Figure 4 A); relative length of fingers II

Published as part of Matsui, Masafumi, Nishikawa, Kanto & Yambun, Paul, 2014, A new Leptolalax from the mountains of Sabah, Borneo (Amphibia, Anura, Megophryidae), pp. 440-452 in Zootaxa 3753 (5) on pages 442-449, DOI: 10.11646/zootaxa.3753.5.3, http://zenodo.org/record/228774