Anacroneuria marlieri Froehlich, 2001 (Figs. 1D, 3 A–F) Anacroneuria marlieri Froehlich, in Ribeiro-Ferreira & Froehlich, 2001: 191; Froehlich, 2003: 130; Ribeiro & Rafael, 2009: 7; Ribeiro et al., 2009: 34; Froehlich, 2010: 161; Ribeiro & Gorayeb, 2014: 18; Menezes et al., 2020: 468. Distribution. BRAZIL: States of Amazonas, Amapá, Maranhão, Mato Grosso, Pará and Roraima. Material examined. BR, MA: Carolina municipality, PARNA Chapada das Mesas, Sucuruiu creek, 07º07’05.6”S, 47º18’31.6”W, 20 – 31/VIII/2013, F. Limeira-de-Oliveira and T.A. Silva, 1 male; same data except 01 – 10/X/2013, J.A. Rafael, F. Limeira-de-Oliveira and T.A. Silva, 1 male; same data except 10 – 20/IX/2013, F. Limeira-de-Oliveira and T. A. Silva, 2 males. Carolina municipality, PARNA Chapada das Mesas, Estiva creek, 07º06’59.8”S, 47º21’21.0”W, 20 – 31/VIII/2013, J.A. Rafael, F. Limeira-de-Oliveira and T. A. Silva, 1 male. Same data except 20 – 30/IX/2013, J.A. Rafael, F. Limeira-de-Oliveira and T.A. Silva, 1 male. Codó municipality, Buriti Corrente creek, 04º45’07”S, 43º39’59”W, 04/X/2019, D.S.S. Araujo, C.L. Franco, J.I.S. Jesus, M.A.G. Almeida, 1 male (reared). Morphometric data. Male (n=7) forewing length: 9 mm. Remarks. Our material has the same pattern of spots on the head, pronotum, and wings as illustrated by Ribeiro & Rafael (2009) (Figs. 3A and 3C). The penial armature of the male seems to vary, with the apex of our specimens in lateral view strongly rounded (Fig. 3F), different from the tapered and acute apex illustrated by Froehlich (2001) and Ribeiro & Rafael (2009). This species is widespread, reported from areas covered by tropical savanna or Cerrado in the states of Maranhão, Mato Grosso and Tocantins; and by Amazon Forest in the states of Amazonas and Roraima (Ribeiro & Santos 2018; Rippel et al. 2019; Menezes et al. 2020) (Fig. 1D). In the State of Maranhão, the species is also widespread, occurring in the East (municipality of Codó), South (municipality of Carolina), and West of the State (municipality of Imperatriz) (Fig. 1D).

Published as part of Carvalho, André De Sousa, Almeida, Lucas Henrique De, Azevêdo, Carlos Augusto Silva De & Lima, Lucas Ramos Costa, 2020, The genus Anacroneuria Klapálek, 1909 (Plecoptera: Perlidae) from Maranhão State, northeastern Brazil, pp. 435-444 in Zootaxa 4860 (3) on page 439, DOI: 10.11646/zootaxa.4860.3.8, http://zenodo.org/record/4414085