Iodes cirrhosa Turcz. Fig. 12.14–12.23 Material examined Specimen used for endocarp and fruit description VIETNAM • “ Indo-chine, Ba Drang ”; 18 May 1921; B. Hayata 672; P[MNHN-P-P06672331]. Other material VIETNAM • “ Indo-chine ”, Annam; 23 May 1923; E. Poilane 6622; P[MNHN-P-P06672332]. Description FRUIT. Elliptical, cleft at the base, red when mature. Epicarp puberulent with yellow simple hairs with granular ornamentation, shriveled when dry, revealing the underlying endocarp ridges. Mesocarp 270– 320 µm thick when dry. Length 12–20 mm, width 10–15 mm, thickness 7–10 mm. ENDOCARP. Cream, elliptical in lateral view, lenticular in transverse section, length ca 12.6 mm, width ca 9.5 mm, thickness ca 4.5 mm. Keel thick, surrounding the endocarp in the plane of symmetry, with the thicker margin containing the primary vascular bundle embedded within the endocarp wall. Apex asymmetrical with an acute protuberance on either side in lateral view; base flat and cleft on one side. Outer surface of the endocarp ridged and faintly rugose, with two thin rectangular to sharp ridges on each face, the median ridge running from the base up to ¾ of the endocarp length and the second ridge running up to the subapical part. Endocarp possessing a symmetrical pair of horn-like protrusions compressed on the keel, positioned eccentrically and subapically one on either endocarp face, each protrusion with a central pit. Endocarp wall 380–405 µm thick excluding ridges (813–1064 µm thick including ridges). Endocarp wall with three cell layers, sometimes homogeneous: outermost layer with 15–17 rows of isodiametric, occasionally anticlinally oriented cells, cells 22–49 µm in diameter, followed by a layer with 3–4 rows of small periclinally oriented cells, cells 7–10 µm in width; innermost layer with one row of periclinally oriented cells, cells ca 6.5 µm in width, covering the inner endocarp surface with regularly spaced, sessile rounded papillae, papillae 8.6–15.5 µm (av. 10.2 µm) in diameter, ca 460 papillae per 0.25 mm 2. Locule surface lacunate. Fig. 12. (see full caption on page 32) Remarks The endocarp was considered “smooth inside” (Sleumer 1971). Here we demonstrate that the locule is densely covered by papillae. These papillae are difficult to observe without SEM.

Published as part of Rio, Cédric Del, Stull, Gregory W. & Franceschi, Dario De, 2020, Survey of the fruits and endocarps of Icacinaceae (Lamiids, Icacinales), pp. 1-130 in European Journal of Taxonomy 645 on pages 34-36, DOI: 10.5852/ejt.2020.645, http://zenodo.org/record/3829651