§ Cyclocephala carbonaria Arrow, 1911 Cyclocephala carbonaria Arrow 1911b: 173 (Nicaragua, Panama) Distribution. Venezuela (EO): Joly 1995: 171 (EO); Moore et al. 2018b: 156 (EO). Other: Arrow 1911b: 173 (Nicaragua, Panama), 1937b: 9 (Ecuador, Nicaragua, Panama); Blackwelder 1944: 251 (Ecuador, Nicaragua, Panama); Endrödi 1966a: 164 (Bolivia, Colombia, Costa Rica, Ecuador, Mexico, Nicaragua, Panama), 1985a: 40, 117 (Bolivia, Costa Rica, Ecuador, Mexico, Nicaragua, Panama); Joly 1995: 171 (Colombia, Costa Rica, Ecuador, Nicaragua, Panama); Ratcliffe 2003: 99 (Costa Rica, Honduras, Mexico, Panama); Moore et al. 2018b: 156 (Belize, Bolivia, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama). Note. Dechambre (1997a) splits the species Cyclocephala carbonaria into a number of species, with C. carbonaria itself being restricted to Central America. According to this, South American specimens previously identified as C. carbonaria, such as those mentioned for Venezuela in Joly (1995: 171) and Moore et al. (2018b: 156) (the latter who simply copied from older publications), must thus belong to other species. Ratcliffe (2003: 99) suggests that more research needs to be done to establish if the South American specimens actually represent separate species or if they are merely intraspecific variations within C. carbonaria. The taxa which are described by Dechambre (1997a) and which are known to occur in the research area are C. boucheri and C. ergastuli. These are considered valid species by us and are included in this checklist.

Published as part of Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, pp. 1-306 in Insecta Mundi 732 (732) on page 165, DOI: 10.5281/zenodo.3678492