10. A. ater (De Geer, 1774) Lentz 1879; Bercio & Folwaczny 1979; Alekseev & Sakhnov 2000; Alekseev 2014. Voucher specimens: 19 [Mel’nikovo, 30 April 1997; Chekhovo, 17 May 1998; 3 km NE Zelenogradsk, 18 June 1998; Cherniakhovsk, 06 May 1999; Mezhdurech’e, 13 June 2000; Zelenogradsk, 13 May 2001; Kolosovka, 02 June 2001, 2 exp.; ibidem, 11 June 2003; Nizinnoe, 12 June 2007, 2 exp.; Krasnoles’e, 08 June 2008; Primorsk, 27 May 2015; Medovoe, 10 april 2018, 2 exp.; Timofeevka, 30 April 2018, 2 exp.; Mechnikov, 07 May 2018, 2 exp.]. Kaliningrad Region: Chern. (Insterburg), Gur. (Juditten); Zel. (Rauschen); Bagr., Chern., Gur., Krasn., Nem., Nest., Pravd., Zel. Comments: Widely distributed and frequent heliophilous species, associated with dung of domestic herbivores.

Published as part of Alekseev, Vitalii I., 2018, Scarabaeoidea (Insecta: Coleoptera) Of The Kaliningrad Region (Russia): The Commented Actual Checklist, Assessment Of Rarity And Notes To Regional Protection, pp. 111-152 in Acta Biologica Universitatis Daugavpiliensis 18 (2) on page 125, DOI: 10.5281/zenodo.10979877