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Mesosciophilidae is one of the extinct families of the suborder Nematocera in Diptera. Six genera and 17 species of mesosciophilids have so far been described from the Holarctic Region, including China, Kazakhstan, Kirghizia, Russia and Transbaikalia. Herein, a new genus with two new species, Similsciophila singularis gen. et sp. nov. and Similsciophila sinuata sp. nov. from the late Middle Jurassic, Jiulongshan Formation of Daohugou Village, Inner Mongolia, China, are described based on their venation and body characters. A new key to genera of mesosciophilid gnats is provided.http://zoobank.org/urn:lsid:zoobank.org:pub:8A772D35-29A8-49AA-9B7F-D9994992EE71
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