Forest sites are formed in the interaction of heterogeneous components of biogeosystem. The increase or decrease of biological component in forest biogeosystems defines the different types of strategy with opposite characteristics, representing the different lines of adaptation (to the biological medium and abiotic environment). The manifestation of adaptive strategy requires appropriate tools, which are evolving on the basis of accumulated property (information capacity) and prevailing forest sites. Forest sites can be seen as an external “order” on the evolution of adaptations. Silvicultural properties of forest biological systems work enough in the interaction of different forms of matter (biological, abiotic), mapping out a hierarchy of biogeosystems. The information potential of forest biota interconnects different levels of systemic organization of forest biogeosystems. Further development of biogeocenology and the theory and ecosystems will allow to investigate the relationship of forest biota with exogenous forest-forming factors.