Abstract The diversification of animal-pollinated angiosperms is related to divergence in floral characteristics promoted by adaptations to different pollinators. According to prevailing evolutionary theory, this macroevolutionary pattern results from adaptive local or regional differentiation of pollination-related features in response to spatial divergence in pollinators. This crucial process links the micro- and macroevolution of floral adaptation, yet it has received much less attention than either floral diversification of species in a phylogenetic context, or pollinator-mediated phenotypic selection on pollination-related traits within populations. This chapter includes two components.