Milviscutulus mangiferae (Green, 1889) (Figs 37, 38) Lecanium mangiferae Green, 1889: 249 Diagnosis. Dorsum with anal plates pyriform, each with 4 apical setae (Figs 37D, 38E); dorsal setae clavate (Fig. 38F); duct tubercles present (Fig. 38D). Marginal setae spinose, mostly with fimbriate apices (Fig. 38M). Venter with multilocular disc-pores usually each with 7–10 loculi, present on posterior area of abdomen, occasionally with a few laterad of metacoxa (Fig. 38K); antenna 8 segmented (Fig. 38P) (partially adopted from Williams & Watson 1990; Hodgson 1994). Material examined. 3 ♀♀, LAOS, Phou Khao Khuay National Bio-Diversity Conservation Area, Thaphabath Dist., Bolikhamsai Prov., 4.v.2015, coll. J.Y. Choi, on Mangifera indica L. (Anacardiaceae); 1 ♀, Vangvieng Dist., Vientiane Prov., 8.ii.2017, coll. P.P. Soysouvanh, on Chrysalidocarpus lutescens H. Wendl. (Arecaceae). Hosts. Polyphagous. According to García Morales et al. (2016), M. mangiferae has been recorded from plants belonging to 82 genera in 42 families. In Laos, it has been found on Dracaena sp. (Asparagaceae) and Ficus sp. (Moraceae) (Suh & Bombay 2015). Distribution. All zoogeographical regions; Oriental Region (India, Indonesia, Laos, Malaysia, Philippines, Singapore, Sri Lanka, Taiwan, Thailand and Vietnam) (Suh & Bombay 2015; García Morales et al. 2016). Economic importance. Milviscutulus mangiferae was listed as a major pest of mango (Mangifera indica) in Israel, U.S.A. and the Pacific region (Gill & Kosztarab 1997), causing yellowing, leaf drop, and death of branches (Grimshaw & Donaldson 2007). Remarks. Milviscutulus mangiferae is closely related to M. spiculatus Williams & Watson, 1990, but is easily differentiated by having clavate or capitate dorsal setae and mostly fimbriate marginal setae; in contrast, M. spiculatus has flagellate to lanceolate but never bluntly clavate or capitate dorsal setae, and the marginal setae are mostly pointed. For comparisons with other species of Milviscutulus, see Williams & Watson (1990).

Published as part of Choi, Jinyeong, Soysouvanh, Pheophanh, Lee, Seunghwan & Hong, Ki-Jeong, 2018, Review of the family Coccidae (Hemiptera: Coccomorpha) in Laos, pp. 1-62 in Zootaxa 4460 (1) on pages 42-44, DOI: 10.11646/zootaXa.4460.1.1, http://zenodo.org/record/1459506