M ore than a century ago, Darwin (lg59) observed that "the number of bumblebees in any district depends in a great measure upon the number of fieldmice, which destroy their combs and nests ... che number of mice is largely Jependent, as everyone knows, on the number of cats .. .it is quite credible that the presence of a feline animal in large numbers in a district might uetermine, through the intervention first of mice and then of bees, the frequency of certain flowers in that district!" (p. -125). Dar\'{in recoglllzed the importance of interactions among organisms, specifically the role that pollinators play as links in communities. Yet, almost 140 years later, our understanding of pollination interactions is still rudimentary. The extent of dependence and linkage in pollination sys-