This paper is devoted to a systematic study and characterizations of the fundamental notions of variational and strong variational convexity for lower semicontinuous functions. While these notions have been quite recently introduced by Rockafellar, the importance of them has been already recognized and documented in finite-dimensional variational analysis and optimization. Here we address general infinite-dimensional settings and derive comprehensive characterizations of both variational and strong variational convexity notions by developing novel techniques, which are essentially different from finite-dimensional counterparts. As a consequence of the obtained characterizations, we establish new quantitative and qualitative relationships between strong variational convexity and tilt stability of local minimizers in appropriate frameworks of Banach spaces.