Uca lactea is a crab belonging to the order Brachyura, but there is no report about its complete mitochondrial genome (mitogenome) in the literature. In this paper, the complete mitogenome of U. lactea was sequenced and the phylogenetic relationship was analyzed based on nucleotide sequences of the 13 protein-coding genes (PCGs) using the maximum likelihood (ML) method. The complete mitogenome of U. lactea is 15,659 bp (GenBank accession No. KY865330) in length and contains 22 tRNA genes, 13 protein-coding genes (PCGs), 2 rRNA genes, and a D-loop region. All PCGs were initiated by ATN codons. The nucleotide composition was also biased toward A + T nucleotides (69.48%). Phylogenetic analysis showed that U. lactea can be placed within the family Ocypodidae. This mitogenome sequence will provide a better understanding for crab evolution in the future.