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Морфологические формы кедра сибирского (Pinus sibirica Du Tour) в высокогорных лесах Северо-Чуйского хребта: 1. Морфологический аспект

Морфологические формы кедра сибирского (Pinus sibirica Du Tour) в высокогорных лесах Северо-Чуйского хребта: 1. Морфологический аспект

Abstract

In the upper forest belt in Altai Mountains, Russia, the coniferous species are the main forest-forming trees. Their living form is straight-stemmed trunk tree. Moving to the climatic lines, individuals of the transition living morphological forms of these species appear in the forest belt. The most diversity of the forms is in the forest-tundra ecotone. We describe the morphological forms of Pinus sibirica Du Tour growing at the upper forest belt in the headwater Aktru, Severo-Chuisky Range, the Altai Mountains. The main living form of Pinus sibirica is tree (straight-stemmed or lean-stemmed trunk, oneor multi-stemmed, oneor multi-top tree) in forest communities here. The basis of its crown is branches of primary crown. The secondary crown do not develop. The transition morphological forms are here: tree or stumps with power candelabra branches, "living" windfallen trees, elfin wood form in the tree butt of the straight-stemmed trunk tree, and shrub-like form. The elbow of the candelabra branch can be produced by both orthotropic growth of the second order branch on first order branch and the elbow of the first order branch. The shrub-like form is formed by an enlargement of the first order branches which are located at the butt of small trees since decline of their top. The elfin wood form grows at the tree butt between buttress flares of the mother tree. The distal part of the form can transform from plagiotropic growth to orthotropic. In fallen trees (if one or some roots and xylem strand are living) some branches change previous growth on orthotropic. All the forms can bear cones for several decades and root if contact with soil. The following conditions and factors promote production of these morphological forms: great insolation, abrupt difference of day and interseasonal temperatures, debris flows, avalanches, stone soil, its grade, firm red heart, shallow root system, and contact of the branches with mellow and moist soil. Such forms often locate in open communities and on glades and at edges in close communities. In one Siberian stone pine individual the combination of the main living form and related with it the morphological structures that are potentially the following vegetative generations can be considered as one of adaptations for sellf-maintenance of the Pinus sibirica populations.

В верхней части лесного пояса Северо-Чуйского хребта Центрального Алтая (горно-ледниковый бассейн Актру) основной жизненной формой кедра сибирского (Pinus sibirica Du Tour) является прямостоячее одноствольное однои многовершинное дерево. Здесь же встречаются другие морфологические формы: (1) деревья и «живые» пни с мощными ветвями-канделябрами, (2) «живой» валежник, (3) кустовидная форма, (4) стланниковидные образования. Их появлению способствуют такие условия и факторы, как высокая интенсивность освещения, резкие перепады суточных и межсезонных температур, сели и лавины, уклон каменистой поверхности, сердцевинные гнили ствола, поверхностная корневая система, соприкосновение ветвей с рыхлым и влажным субстратом. Совмещение у кедра сибирского в пределах одной особи основной жизненной формы и связанных с ней структур, которые потенциально являются последующими вегетативными поколениями, может рассматриваться в качестве одной из адаптаций для самоподдержания его популяций.

Keywords

МОРФОЛОГИЧЕСКИЕ ФОРМЫ, ВЕРХНЯЯ ЧАСТЬ ГОРНО-ЛЕСНОГО ПОЯСА, СЕВЕРО-ЧУЙСКИЙ ХРЕБЕТ, ЦЕНТРАЛЬНЫЙ АЛТАЙ, КЕДР СИБИРСКИЙ

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selected citations
These citations are derived from selected sources.
This is an alternative to the "Influence" indicator, which also reflects the overall/total impact of an article in the research community at large, based on the underlying citation network (diachronically).
BIP!Citations provided by BIP!
popularity
This indicator reflects the "current" impact/attention (the "hype") of an article in the research community at large, based on the underlying citation network.
BIP!Popularity provided by BIP!
influence
This indicator reflects the overall/total impact of an article in the research community at large, based on the underlying citation network (diachronically).
BIP!Influence provided by BIP!
impulse
This indicator reflects the initial momentum of an article directly after its publication, based on the underlying citation network.
BIP!Impulse provided by BIP!
0
Average
Average
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