Figure 8. Behavioral sequence of the hermit crab Pagurodofleinia doederleini transferring the sea anemone Stylobates calcifer sp. nov. from the original snail Glossaulax reiniana shell to a new one. (A) The hermit crab begins tapping the pedal disk and column of the sea anemone with its walking legs and chelipeds. (B) The hermit crab uses one or both of its chelipeds to pinch and remove S. calcifer sp. nov. from the G. reiniana shell. (C) Stylobates calcifer sp. nov. displays no shell-mounting action after being removed from the original shell. (D) The sea anemone is turned to the upside-down position (facing up the pedal disk) by the crab. (C) The hermit crab rides on the pedal disk of the sea anemone and fits the shell to the curved line of the pedal disk. (E) The sea anemone settles down in a new shell of the host hermit crab with a usual position.

Published as part of Yoshikawa, Akihiro, Izumi, Takato, Moritaki, Takeya, Kimura, Taeko & Yanagi, Kensuke, 2022, Carcinoecium-Forming Sea Anemone Stylobates calcifer sp. nov. (Cnidaria, Actiniaria, Actiniidae) from the Japanese Deep-Sea Floor: A Taxonomical Description with Its Ecological Observations, pp. 127-152 in The Biological Bulletin 242 (2) on page 139, DOI: 10.1086/719160, http://zenodo.org/record/7573707