The species includes 39 subspecies. The type locality is Georgia. Seven populations of D. (C.) s. scabricolle are known from the republic. One population of the type subspecies is situated in north Armenia. D. (C.) s. shirakense, ssp. n. (type locality Lernavan environs, 40°48΄10.57΄΄N 44°09΄58.10΄΄E, 1893 m) is distributed all over west part of Armenia and in north-east Turkey. Seven more subspecies are recognized in Armenia: D. (C.) s. tavushense, ssp. n. (type locality – Idzhevan environs, Acharkut, about 41°2΄9΄΄N, 45°5΄6΄΄E, 800 m) in north Armenia, D. (C.) s. gegarkunicum, ssp. n. (type locality – Akhberk environs, 40°32΄58΄΄N, 45°17΄09΄΄E, 2280 m) around north part of Sevan Lake, D. (C.) s. sevangense Reitter, 1889 (type locality – environs of Sevan-city ) - high mountains all acrros cenral part of Armenia, D. (C.) s. pseudosevangense, ssp. n. (type locality – Noravan environs, 39°32΄15΄΄N, 46°07΄56΄΄E, 2000 m) at the east part of D. (C.) s. sevangense area, D. (C.) s. vaykense, ssp. n. (type locality – Vayk environs, 3.6 km northwards Zaritap, 39°40΄11΄΄N, 45°30΄28΄΄E, 1300 m) – south of Marz Vayots Dzor, D. (C.) s. tekhense, ssp. n. (type locality – Tekh, 39°34΄6΄΄N, 46°25΄52΄΄E, 1600 m) – north-east of Marz Syunik, D. (C.) s. tuzovi, ssp. n. (type locality – Megri environs) – south-east part of Armenia with penetration to Nakhchivan Autonomous Republic; a new subspecies is described from Nagorno-Karabakh Republic: D. (C.) s. artsakhense, ssp. n. (type locality – Shishkend environs, 39°45΄52΄΄N, 46°48΄09΄΄E, 1190 m). Six subspecies are accepted for Nakhchivan Autonomous Republic including D. (C.) s. buzgoviense, ssp. n. (type locality – Buzgov environs, about 39°31΄7΄΄N, 45°23΄23΄΄E, 1700 m), D. (C.) s. shakhbuzum, ssp. n. (type locality – Shakhbuz District, Bichenek environs, 39°30΄50΄΄N, 45°46΄28΄΄E, 1940 m), D. (C.) s. araxense, ssp. n. (type locality – Negram environs, 39°5΄N, 45°24΄13΄΄E, 790 m), D. (C.) s. paiz Danilevsky, 1999 (type locality – Paiz env., 39°25΄30΄΄N, 45°23΄14΄΄E, 1300 m), D. (C.) s. nakhiczevanum Danilevsky, 1999 (type locality - south slope of Bichenek pass, 39°31΄58΄΄N, 45°47΄51΄΄E, 2100 m), D. (C.) s. dobrovljanskii Suvorov, 1915 (type locality – Kyukyu village, about 39°31΄27΄΄N, 45°37΄18΄΄E). Two subspecies are distributed in Azerbaijan: D. (C.) s. modestum Tournier, 1872, nom. rest. (type locality – Azerbaijan, Shamkir [before Annenfeld], 40°50΄83΄΄N, 46°01΄02΄΄E) – from north-west Azerbaijan and D. (C.) s. corpulentum Ménétriés, 1832 from Talysh mountains (including Iranian part). Six subspecies are distributed in Iran: D. (C.) s. babakkalense, ssp. n. (type locality – Āzarbāijān-e Sharqi province, Babak-Kale environs, 38°50΄48.48΄΄N, 46°59΄2.40΄΄E, 2100 m) – Karadag Ridge in north-west Iran, D. (C.) s. subcorpulentum Breuning, 1946 (type locality – Khoy [about 38°33΄N, 44°57΄E]) – eastwards Urmia lake, D. (C.) s. sagezense Lazarev, 2013 (type locality – Kordestan, 6 km N Sagez) east- and southwards Urmia lake, D. (C.) s. sahandum, ssp. n. (type locality – Āzarbāijān-e Sharqi province, Kandovan environs, southwards Tabriz, 37°48΄21.96΄΄N, 46°18΄27.72΄΄E, 2350 m) – Sahand Mts. environs and eastwards, D. (C.) s. skoupyi Lazarev, 2013 (type locality – eastwards Khalkhal, 37°35΄ N 48°38΄E, 2068 m) – Ardabil province, D. (C.) s. gilanense Lazarev, 2013 (type locality – Gilan, Kharpu env.) – mountains south-westwards Caspean see. Sixteen subspecies are distributed in Turkey: D. (C.) s. crassofasciatum Özdikmen, 2013 (type locality – Balikesir province, Edremit, Sarikiz hill, 39°42΄N, 26°49΄E”) – the most north-western area of Turkey, D. (C.) s. balikesirense Breuning, 1962 (type locality – Balikesir environs [about 39°38΄N, 27°52΄E]) – central part of Balikesir prov., D. (C.) s. uludaghicum Breuning, 1970 (type locality – Ulu-Dagh Mountain near Bursa [about 40°06΄N, 29°10΄E]) – Bursa province, D. (C.) s. inonuense, ssp. n. (type locality – northwards Inönü westwards Eskişehir, about 39°48΄N, 30°08΄E) – Eskişehir province, D. (C.) s. apakoyense, ssp. n. (type locality – Apaköy environs southwards Denizli, about 37°30΄29΄΄N, 29°21΄19΄΄E) – Denizli province, D. (C.) s. paphlagonicum Breuning, 1962 (type locality – Kastamonu env. [about 41°23΄N, 33°46΄E].) – Kastamonu, Çankiri, Ankara and probably Çorum provinces, D. (C.) s. yahyaliense Bernhauer & Peks, 2011 (type locality – Kayseri province, 38 km eastwards Yahyali (120 km eastwards Niğde) [about 38°5΄24΄΄N, 35°45΄34΄΄E] 1900 m.) – Kayseri, Niğde and Adana provinces, D. (C.) s. antonkozlovi Danilevsky, 2017 (type locality – Güney village environs, 37°36΄57΄΄N, 34°29΄50΄΄E) – Niğde province, D. (C.) s. caramanicum K. Daniel & J. Daniel, 1903 (type locality – Bulghar Maaden [about 37°27΄N, 34°37΄26΄΄E]) – Niğde province, D. (C.) s. salhanum, ssp. n. (type locality - Bingöl province, 6 km south-eastwards Salhan, Buglan Pass, 38°56΄9΄΄N, 41°6΄42΄΄E, 1800 m) – from Kahramanmaraş to Bitlis, D. (C.) s. alucranum, ssp. n. (type locality – Alucra environs, about 40°19΄7΄΄N, 38°45΄56΄΄E) – Girezun province, D. (C.) s. lazistanum, ssp. n. (type locality – Gümüşhane environs, 40°27΄26΄΄N, 39°28΄14΄΄E, 1300 m) – Gümüşhane province; D. (C.) s. hajdajorum, ssp. n. (type locality - Başören environs [about 38°28΄N, 30°27΄E], 1800-2000 m) – Afyonkarahisar province, D. (C.) s. korbianum, ssp. n. (type locality - Akşehir environs [about 38°21΄N, 31°25΄E]) – Konya province; D. (C.) s. gazii, ssp. n. (type locality – Seyitgazi environs [about 39°26΄44΄΄N 30°41΄38΄΄E]) – Eskişehir province.