The “ browni group” The wing patterns of all species of this group are very simple and consistent. The dorsal wing surface in males is structurally colored blue or purple, while the ventral wing surface is brown or reddish brown with simple and faint patterns of medial and submarginal bands. These subtle pattern elements proved to be among the most useful to distinguish species within the group. Secondary male wing characters, such as the shape and coloration of the androconial cluster, is also taxonomically useful. The male valvae shapes, although also simple in structure, may differ significantly and consistently between species that are externally almost indistinguishable, like the species-pair P. browni and P. saraha. Since, the entire view of the whole male genitalia doesn’t provide sufficient information to discriminate species, we prefer to illustrate only the valvae in lateral view. The species of this presumably monophyletic group share the following combination of morphological characters which seem to be unique in Penaincisalia sensu Robbins (See Table 1): 1-Hind wing tornus in males appears as a spatulate and bulbous lobe (Figure 1B). 2-Sexual dimorphism in wing shape, where the females have a long tail from the CuA2 vein, accompanied by a small lateral lobe in the hindwing tornus (Figure 1B, C). 3-Male androconial cluster comprised by a scent pad in the apical part of the discal cell and a minute scent patch situated distally at the erection of lower discocellular vein (Figs 26-33). The scales in the scent pad and scent patch are arranged tightly and oriented almost perpendicularly to the wing membrane; they are identical in size (100 µ in length and 30 µ in width) and nanostructures (Bálint et al., in prep.). 4-The male genital valva possesses a prominent, short triangular process at the ventral base (Figure 34– 43). 5-Female genital posterior portion of ductus bursae shorter than anterior portion of ductus bursae (Figures 44–49; anterior and posterior portion defined in fig. 44). Characters: 1 = Hindwing tornus in males appears as a spatulate and bulbous lobe. 2 = Sexual dimorphism in wing shape, where the female has a long tail from the CuA2 vein, accompanied for a small lateral lobe in the hindwing anal corner. 3 = Male androconial cluster comprised by a scent pad in the apical part of the discal cell and a minute scent patch situated distally at the erection of lower discocellular vein. 4 = male genital valvae possesses a prominent, short triangular ventral process in the ventral edge. 5 = female genital ductus bursae posterior portion is shorter than the anterior one.

Published as part of Prieto, Carlos, Bálint, Zsolt, Boyer, Pierre & Micó, Estefanía, 2008, A review of the " browni group " of Penaincisalia with notes on their distribution and variability (Lepidoptera: Lycaenidae: Eumaeini), pp. 1-24 in Zootaxa 1941 (1) on pages 3-5, DOI: 10.11646/zootaxa.1941.1.1, http://zenodo.org/record/5231015