Key to species of genus Brithura 1. Abdominal tergites with big blackish brown spots at lateral margins .......................................................................... 2 - Abdominal tergites without big spots at lateral margins ............................................................................................. 3 2. Wing dark grayish brown with some light golden areas; male hypopygium with outer gonostylus thick, laterally with three extensions at tip, medially with pointed spine at bottom (Fig. 12) .......................................... B. aureola sp. nov. - Wing yellowish; male hypopygium with outer spine of outer gonostylus conspicuous (Fig. 8) ............ B. shii sp. nov. 3. All femora with conspicuous subterminal ring............................................................................................................. 4 - Posterior femur without subterminal ring................................................................................................................... 10 4. Femora with tips yellow, subterminal ring brown or black .......................................................................................... 5 - Femora with tips brown or black, subterminal ring yellow.......................................................................................... 6 5. Femora brownish yellow, apical ring yellow, preceded by a narrow black ring; outer gonostylus with two dissimilar spines at middle outer side (Edwards 1916, p264, Pl. XII. fig. 12) ........................................ B. crassa Edwards, 1916 - Femora light brown with a narrow virtually terminal yellow ring, preceded by a subequal brown ring; outer gonosty- lus not as above (Alexander 1964, p87, Pl. I. fig. 29) ................................................ B. nielseniana (Alexander, 1964) 6. Abdominal tergites dark brown ................................................................................................................................... 7 - Abdominal tergites not as above .................................................................................................................................. 9 7. Scutellum obscure yellow; pleura uniform in color except by pale dorsal surface of pleurotergal tubercle; outer spine of outer gonostylus large and conspicuous (Alexander 1935, p90, Pl. II. fig. 25) .... B. fracticosta (Alexander, 1935) - Scutellum dusky brown; pleura with variegated with silvery, additional to pale pleurotergal tubercle; outer gonosty- lus not as above ............................................................................................................................................................. 8 8. Antennal flagellum brownish yellow; femora yellowish brown, with base yellow; outer gonostylus with a short slen- der rod at outer margin (Alexander 1929, p317, Pl II. fig. 19) ............................................ B. sancta Alexander, 1929 - Antennal flagellum uniformly dark brown; femora yellow; outer spine of outer gonostylus small and thin (Alexan- der 1935, Pl. II. fig. 26) .............................................................................................. B. fractistigma Alexander, 1925 9. Abdominal tergites chestnut brown?strips of prescutum disintegrated into two lines of subrectangular brown spots; outer gonostylus laterally with three processes (Fig. 20) ................................................................ B. stigmosa sp. nov. - Abdominal tergites bright orange; strips of prescutum continuous; outer gonostylus outer gonostylus at near apex with a cylindrical finger-like lobe on outer margin (Alexander 1971, p378, fig. 15). B. keiliniana (Alexander, 1971) 10. Fore and middle femora with conspicuous subterminal ring...................................................................................... 11 - Femora without subterminal ring................................................................................................................................ 13 11. Ground color of abdomen light brownish orange; outer gonostylus apically bent forward greatly with a short subtri- angular posterior process (Liu & Yang 2009, p54, fig. 5) ........................................ B. jinpingensis Liu & Yang, 2009 - Ground color of abdomen dark brown or brownish black.......................................................................................... 12 12. Femora yellow; outer gonostylus anteriorly with a long slender rod (Alexander 1971, p378, fig. 16) ........................ ...................................................................................................................................... B. brulleana (Alexander, 1971) - Femora reddish-brown; outer gonostylus slightly bent forward, basal outer margin with an erect slender rod (Edwards 1916, p263, Pl. XII. fig. 10) ........................................................................... B. imperfecta (Brunetti, 1913) 13. Tip of vertical tubercle polished, light yellow; abdominal tergites uniformly orange, outer lateral angles not pale .... ................................................................................................................................... B. argyrospila (Alexander, 1935) - Vertical tubercle uniformly darkened; color of abdominal tergites not as above, outer lateral angles of basal abdomi- nal tergites pale ........................................................................................................................................................... 14 14. Abdominal tergites cinnamon brown; outer gonostylus with two processes terminating in long slender rod .............. ......................................................................................................................................... B. nymphica Alexander, 1927 - Abdominal tergites orange; outer gonostylus not as above ........................................................................................ 15 15. Mediotergite brown with posterior margin yellow; outer gonostylus bent forward slightly at tip, laterally with a con- spicuous spine (Fig. 16) .......................................................................................................... B. flaviflagellum sp. nov. - Mediotergite yellow with posterior margin yellowish brown; outer gonostylus with two lobe-like processes (Liu & Yang 2009, p55, fig. 9) ............................................................................................. B. guangxiensis Liu & Yang, 2009

Published as part of Liu, Qifei & Yang, Ding, 2010, Four new species of the genus Brithura Edwards from China, with an updated key to world species (Diptera, Tipulidae), pp. 52-60 in Zootaxa 2401 on pages 52-53, DOI: 10.5281/zenodo.194087