Euptychia cesarense viloriai Andrade, Pulido, Peña & Lamas, new subspecies (Figs. 2, 6) Euptychia [n. sp.]: Lamas, 2004 Description. MALE, FW length: 17 mm (n=4) (Fig. 6 A). Similar to nominotypical subspecies but DHW with an additional white-pupilled faint black ocellus in Cu2-2A, the lines below are narrower and the ventral ground color is paler (R194, G175, B86), in VFW the ocellus between R5 and M2 is smaller and the dark yellow patches between postdiscal and submarginal lines in M3-Cu1 and Cu1-Cu2 are nearly inconspicuous, in VHW the anal area between the inner margin and the postdiscal line is dusted with white scales. Ventrally, the discal, postdiscal, marginal and submarginal lines are thinner than those of nominotypical subspecies and submarginal line in both wings is more undulated in M2–Cu1. Genitalia as in E. cesarense cesarense. FEMALE, FW length: 17.2 mm (n=7) (Fig. 6 B): Ground color in both wings paler and more yellowish than in E. cesarense cesarense. DFW with a white pupilled black ocellus in M1-M2. DHW with white-pupilled black ocelli in M1-M2, Cu1-Cu2, and a faint one in Cu2-2A. Ventrally, the lines are wider than in E. c. cesarense, and the VFW patch between postdiscal and submarginal lines is more conspicuous. Genitalia as in E. cesarense cesarense. Early stages and host plants. Unknown. Etymology. This subspecies is named after our friend and colleague Angel Viloria, in recognition of his important work on Andean Lepidoptera. Type locality. Perú, Junín, Mina Pichita, 11º05’ S, 75º25’ W, 2100 m. HOLOTYPE: Male, with the following labels: PERU, Junín, 1 Km S Mina Pichita, 11º05’ S, 75º25’W, 2100 m, 21 August 2003, C. Peña Leg., MUSM-ENT-003934, GENITALIA #HWPB-01. ALLOTYPE: Female, with the following labels: PERU, Junín, 1 Km S Mina Pichita, 2100 m, 22 August 2003, C. Peña Leg., MUSM. PARATYPES: PERU, Junín, 1 Km S Mina Pichita, 11º 05’S, 75º 25’W, 2100 m, 12 November 2003, C. Peña Leg, MUSM, DNA Sample 04-55 Carlos Peña (1 male); Junín, 1 Km S Mina Pichita, 11º 05’S, 75º 25’W, 2100 m, 22 August 2003, C. Peña Leg, MUSM (2 females); Junín, 1 Km S Mina Pichita, 11º 05’S, 75º 25’ W, 2100 m, 9 September 2002, J. Grados Leg, MUSM (1 male); Junín, 1 Km S Mina Pichita, 11º 05’ S, 75º 25’ W, 2100 m, 9 August 2002, J. Grados Leg, MUSM (1 female); Junín, 1 Km S Mina Pichita, 11º 05’ S, 75º 25’ W, 2100 m, 13 August 2002, C. Peña Leg, MUSM (2 females); Junín, 1 Km S Mina Pichita, 11º 05’ S, 75º 25’ W, 2100 m, 21 August 2003, J.J. Ramirez Leg, MUSM, Genitalia HWPB-03, MUSM-ENT-003937 (1 male); Junín, 1-3 Km S Mina Pichita, 2100 m, 24 August 1988, G. Lamas Leg, MUSM (1 female); Junín, 1 Km S Mina Pichita, 11º 05’ S, 75º 25’ W, 2100 m, 12 August 2002, A. Guanllo Leg, MUSM (1 female); Junín, 1-3 Km S Mina Pichita, Hda. Naranjal, 2100 m, 17 October 1989, G. Lamas Leg, MUSM (1 female); Junín, 0-1 Km E, Mina Pichita, Hda. Naranjal 2000 m, 18 November 1984, G. Lamas & J.E Pérez Leg, MUSM (1 female) ADDITIONAL MATERIAL: PERU, Pasco, 18 Km S Pozuzo, 10º 16’ S, 75º 33’ W, 1200 m, 29 November 1997, G. Lamas Leg, MUSM (1 male); Cuzco, Cerca Santa Isabel, Río Cosñipata, 1200 – 1500 m, 5-11 February 1975, G. Lamas leg., MUSM-ENT-003941 (1 female); Cuzco, Río Cosñipata, Qbda. Quitacalzón, 1050m, 1-3 Septiembre 1989, G. Lamas Leg. (1 female); ECUADOR, Tungurahua, Río Blanco, 01º23’ S, 78º20’ W, 1000 m, 0 7 July 1966, S.E. Velástegui Leg, MUSM (1 female); Río Machay, 1º 23.75’ S, 78º 16.30’ W, 1700 m, 4-5 July 1993, J. Hall, leg, KWJH (1 male); Río Machay, 1º 23.75’ S, 78º 16.30’ W, 1700 m, 19-20 August 1993, K. Willmott & J. Hall, leg, KWJH (1 female); Río Machay, 1º 23.75’ S, 78º 16.30’ W, 1700 m, 19-20 August 1993, K. Willmott & J. Hall, leg, MECN (1 female). Diagnosis. Euptychia cesarense viloriai males are distinguished from the nominotypical subspecies by having a pupilled ocellus between Cu2-2A of DHW, the VHW anal area dusted with white scales, ventral ground color paler, ventral lines narrower, VHW discal line more oblique and submarginal line in the VFW apex in zigzag, rather different from the smoothly roundend line in the cesarense. Females can be separated by a general paler color, a pupilled ocellus in cell Cu2-2A of DHW and wider VW submarginal lines are evident. Distribution and ecology. Known from Junín, Pasco and Cuzco in Peru. The subspecies flies from 1200 to 2100 m. The existence of the species in several localities of Peru, and northern Colombia, suggests that Euptychia cesarense is a rare but widely distributed species in the Andes, with additional subspecies likely to be discovered in regions where it is currently unknown (Fig. 1). We compared additional material from Ecuador sharing features with cesarense and viloriai, which seems to be a different subspecies, but the status of these spcimen must be reviewed.

Published as part of Pulido-B, Hannier W., Andrade-C, Gonzalo, Peña, Carlos & Lamas, Gerardo, 2011, Two new taxa of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the Andes of Colombia and Peru, pp. 43-51 in Zootaxa 2906 on pages 49-51, DOI: 10.5281/zenodo.277759