Genus Ypsiloncyphon Klausnitzer, 2009 Males of several “ Cyphon -like” Scirtidae whose invaginated abdominal segments 8 and 9, and whose genitalia include Y-shaped components, stood out from other “ Cyphon -like” taxa as a separate entity which Yoshitomi (2005) called the chlorizans -group. Ruta (2007) recognized 3 subgroups in it. Klausnitzer (2009) established the genus Ypsiloncyphon for the chlorizans -group, with the same three species groups. Each of these groups appears to be a monophylum. In contrast, the monophyly of Ypsiloncyphon as a whole is less convincingly documented. Species groups 1 and 2 are Oriental and unknown in the Australasian region where species group 3 is endemic. Species group 3 is very distinct by the male terminalia and females uniformly possess a dictyon in the genitalia which was never before described or illustrated. However, this is apparently a universal feature of the genus, known also in Oriental, Neotropical, and Afrotropical species (R. Ruta, personal communication). Therefore I decided to wait for additional information on the other two species groups before creating a new genus-group name for group 3, which may be required in the future. A general description applicable to all species in group 3 is provided. Previously included species are redecribed from types, and several new species from Australia and New Guinea are named and described.

Published as part of Zwick, Peter, 2014, Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer, pp. 1-41 in Zootaxa 3846 (1) on page 16, DOI: 10.11646/zootaxa.3846.1.1, http://zenodo.org/record/286792