Incoltorrida galoko, new species Figs. 13 (habitus); 36 (map); 40 (habitat) Type Material. Holotype (female): Antsiranana, “Anjiabe Ambony: Ambilobe: Antsabe stairways-like cascade with vertical (!) steps, exposed, extremely hot day, N: -13.60930 E: 48.72120, elev. 303 m, 23 xi 2004, Balke et al. (P25 MD16)” // DNA extraction, # BMNH 670736 (BMNH). Differential Diagnosis. An abundantly distinct species, differentiated from other members of the genus by the combination of small size, broadly oval body form, extreme pronotal sculpture, and each elytron with five costae. Currently know only from the female holotype, which is distinct morphologically, and also distinct according to DNA sequence data published by Monaghan et al. (2009) (Genbank Accession Nos. FJ819704 (partial COI) and FJ818160 (partial 28S)) from the two coexisting species I. magna and I. quintacostata. Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.74/1.14; head width 0.36; pronotum 0.43/0.76; elytra 1.07 /1.14. Dorsum dark brown, venter brown to reddish brown, legs brown to reddish brown except tarsi and femoral-tibial articulations black. Body form very broad, oval (Fig. 13). Sides of frontoclypeal shield very slightly arcuate, narrowest, but only slightly so, at frontoclypeal suture. Head slightly concave in front of eyes. Pronotal sculpture extreme (Fig. 13), ridges distinctly raised and depressions deep; anterior part with three carinae, midlongitudinal carina and oblique carinae on each side. Midlongitudinal carina terminating where, on each side is transverse timidity. Together, sculptural elements on disc somewhat resemble a snarling lion’s face. Each elytron with only five distinct well separated, granulate costae: #1, #2, #3, #5, and #8; costae #4, #6, and #7 absent. Merest hint of two transverse ridges, one linking costae #3 and #5 and one linking costae #5 and #8, later located just behind the humeral umbo. Nine rows of small serial punctures, two rows each between costae #4 and #5, #5 and #6, #6 and margin; three other rows single. Midventral area of metaventral tabella with distinct, widely spaced transverse grooves. Midlongitudinal carina of first abdominal ventrite, strong, extends length of ventrite. Etymology. Named in reference to the type locality. Remarks. The holotype female, which has been extracted for DNA, is unfortunately in pieces, and is missing several leg parts. The parts were glued together for the habitus figures (Fig. 13).

Published as part of Perkins, Philip D. & Bergsten, Johannes, 2019, New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae), pp. 57-96 in Zootaxa 4657 (1) on page 81, DOI: 10.11646/zootaxa.4657.1.2, http://zenodo.org/record/3369801