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Published as part of Duarte, Paulo R. M., Grossi, Paschoal C. & Dupuis, Fabien, 2022, A taxonomic revision of the Bothynus villiersi Endr��di, 1968 species group (Coleoptera: Scarabaeidae: Dynastinae), pp. 49-66 in Zootaxa 5093 (1) on page 61, DOI: 10.11646/zootaxa.5093.1.3, http://zenodo.org/record/5901043
FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endr��di, 1968; B, Bothynus horridus Endr��di, 1968; C, Bothynus alvarengai Mart��nez, 1983; D, Bothynus belemensis new species.
Coleoptera, Insecta, Arthropoda, Dynastidae, Animalia, Biodiversity, Bothynus, Taxonomy
Coleoptera, Insecta, Arthropoda, Dynastidae, Animalia, Biodiversity, Bothynus, Taxonomy
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