Aegidium onorei Frolov, Akhmetova and Vaz-de-Mello, sp. nov. (Figure 4 (a–j)) Type locality Ecuador, Cotopaxi, San Francisco de las Pampas. Type material examined Holotype, male at CEMT labeled ‘ ECUADOR: Cotopaxi. San Fco de las Pampas. III-1983, GOnore’ (Figure 4 (b)). Paratypes: ECUADOR. One male at MNHN labeled ‘ Ecuador’; one male at CEMT labeled ‘ Ecuador A. Louste leg’. Cotopaxi: one male at BGCO labeled ‘ Ecuador Cotopaxi Las Pampas Dec. 1989 Legit: G. Onore’; one male and two females at CEMT labeled ‘ ECUADOR: Cotopaxi. San Fco de las Pampas. III-1983, GOnore’. Pichincha: one male at CMN labeled ‘ Ecuador Santo Domingo IV-82 G. Onore’; one male at CEMT labeled ‘ ECUADOR Sto. Domingo IV.82. G. Onore leg.’; one male and two females at MHNG labeled ‘ Équateur Pichincha Santo Domingo de los Colourados 500 m G. Onore IV.83 ’. Diagnosis Aegidium onorei sp. nov. can be separated from other Aegidium species by the presence of granulate sides of the pronotum and mesotibiae with a tuft of dense setae on the inner side apically in males (Figure 4 (a,c)). Description Holotype, male (Figure 4 (a,e–i)). Body length 18.0 mm. Colour uniformly black. Anterior margin of frontoclypeus slightly convex at middle, slightly sinuate each side of medial convexity, rounded laterally, without border. Frontoclypeus punctate with large, rounded punctures separated by about 1 puncture diameter; punctures on anterior part have slightly raised margin giving surface a rugose appearance. Eyes relatively small: width about 1/8 distance between eyes in dorsal view. Pronotum wider than elytra, 1.55 times wider than length, 0.6 times length of elytra. Posterior angles rounded. Anterior margin not bordered. Base of pronotum without furrow, not bordered. Pronotal disc flattened and excavated, with longitudinal depressed line basally. Lateral pronotal processes long, horn-shaped in lateral view, distinctly protruding past lateral margin outline of pronotum in dorsal view, slightly curved towards pronotal centre. Anterior pronotal process cone-shaped. Disc and anterior part of pronotum covered with round to elongate punctures with raised margin; base and lateral sides granulate. Scutellum narrow subtriangular, about 1/11 length of elytra. Elytra 1.2 times longer than wide, with humeral and apical umbones. Elytra widest in basal 1/3, tapering apically in dorsal view. First elytral carina feebly distinct, second indistinct. Elytra covered with relatively dense, round to elongated punctures. Punctures on disc and base have slightly raised margin. Macropterous. Mesotibiae with a tuft of setae ventroapically. Inner apical spur of mesotibia shorter than basal mesotarsomere and not curved downwards (Figure 4 (c)). Aedeagus with relatively short (0.4 times length of phallobase), tapering parameres (Figure 4 (g)). Parameres slightly wider than apical part of phallobase in dorsal view (Figure 4 (h)). Separate ventrobasal transverse plate of parameres absent (Figure 4 (i)). Lateral teeth of parameres not protruding past paramere outline in dorsal view (Figure 4 (h)). Endophallus without strongly sclerotised sclerites (Figure 4 (e)). Cranial part of spiculum gastrale subtriangular, widened and rounded apically (Figure 4 (f)). Female differs from male in having a protibial spur, relatively smaller, punctate pronotum without armature and excavations, and inner protibial tooth absent (Figure 4 (d)). Body length 13–16 mm. Variation Body length of the male paratypes 16.5–22.0 mm. Some males have a relatively smaller pronotum (as wide as or slightly narrower than elytra) with less developed lateral processes. Distribution Aegidium onorei sp. nov. is known from a few mid-altitude localities in western Ecuadorian Andes (Figure 4 (j)).

Published as part of Frolov, Andrey V., Akhmetova, Lilia A. & Vaz-de-Mello, Fernando Z., 2017, Revision of the mainland species of the Neotropical genus Aegidium Westwood (Coleoptera: Scarabaeidae: Orphninae), pp. 1035-1090 in Journal of Natural History 51 (19 - 20) on pages 1063-1065, DOI: 10.1080/00222933.2017.1319519, http://zenodo.org/record/4746652