Sophiodela japonica (Thunberg, 1781), comb. nov. (Figs 7B, 12B) Cicindela japonica Thunberg, 1781: 25. Cicindela chinensis japonica: Fleutiaux, 1892:116. Cicindela chinensis japonica konoi Goto & Iga, 1950:14 [synonymy by Nakane 1976: 4]. Cicindela chinensis yakushimana Nakane, 1963:3 [synonymy by Nakane 1976: 4]. Cicindela (Sophiodela) japonica: Nakane 1955: 26. Cicindela (Sophiodela) chinensis japonica: Cassola & Nidek 1984:11. Cicindela (Sophiodela) japonica: Fukuda et al. 2015:271. Specimens examined. (N = 4) [Japan] 2♂♂ (EUMJ), Fruit Tree Research Center, Shimoidai, Matsuyama-shi, Ehime Pref., 2. X. 2018, R. Ogawa leg; 2♂♂ (EUMJ), ditto, 25. X. 2018, R. Ogawa leg. Description. Internal sac everted dorsally in lateral view. DA largest in the bladders: DLL the second largest: VA as large as B. VLR, BLR and VB not exist. VA short triangular everted dorso-apically. DA everted basally. B oval swollen ventrally, with well sclerotized spines, which are forming a narrow band. DLL swollen dorsally, covered numerously with well sclerotized spines, which are a forming band. Flagellum well short, curved and formed S shape. Medial tooth short, orthogonally curved right ventro-laterally. Remarks. This species shares the characteristics of the position of each bladder with Sophiodela okinawana, S. ferriei, and S. c. chinensis, but is distinguished from the three species by the following characteristics: B oval swollen ventrally; VA triangular everted dorso-apically; medial tooth orthogonally curved.

Published as part of Fukuda, Yuki, Ogawa, Ryo & Hori, Michio, 2019, The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia, pp. 271-308 in Zootaxa 4661 (2) on page 284, DOI: 10.11646/zootaxa.4661.2.3, http://zenodo.org/record/3998528