Philothalpus anapodos Chatzimanolis, new species (Figs. 1, 3–4, 6–7, 11) urn:lsid:zoobank.org:act: 35B1590C-6AF4-4DD8-9AF7-20FF977C3EB4 Type material. Holotype, here designated, male, with labels: “ PERU, Madre de Dios Dept. CICRA field station, ~ 2 km NW of cafeteria, research plot, 12.55212°S 70.10921°W, 295 m, 11–13.vi.2011, Chaboo team, flight intercept trap, PER-11-FIT-023” / “[barcode label] SEMC0985242 ”. In the collection of MUSM. Paratypes. Four:same label as holotype except 9–11.vi.2011, PER-11-FIT-022and barcode label SEMC0955098 (1 ♀ SEMC); same label as holotype except 7–9.vi.2011, PER-11-FIT-021 and barcode labels SEMC0956709, SEMC0956710 (1 ♀ MUSM, 1 ♀ UTCI); same label as holotype except 12.55261°S 70.11008°W, 11–13.vii.2010, PER-10-07-FIT-004, barcode label SEMC0955348 (1 ♀ SEMC). All paratypes with label “ PARATYPE Philothalpus anapodos Chatzimanolis, des. Chatzimanolis 2019 ”. Diagnosis. Among Philothalpus species with glossy integument of head and pronotum, P. chotaenus, P. ecuadorensis, P. juanecoi, P. porphyros, P. portokalis, P. loksos, P. stravos, and P. anapodos all have large, numerous, contiguous to virtually contiguous punctures on pronotum. Philothalpus anapodos and P. portokalis are the only two species in that group that have reddish-brown to orange head and pronotum and punctures on pronotum that are round and not elongated longitudinally. In P. portokalis the head and pronotum are bright orange, the pronotum is strongly concave posteriorly and the apex of the paramere is emarginate, while in P. anapodos the head and pronotum are reddish-brown, the pronotum is weakly concave posteriorly and the apex of the paramere is rounded; P. anapodos also does not have any peg setae on the paramere. Description. Forebody length 5.9–6.7 mm. Coloration of head and pronotum shinning reddish brown; legs, mouthparts, antennomeres 1–3 reddish brown; antennomeres 4–11 brown; elytra shinning metallic purple-brown or green, with golden overtones; elytra epipleuron lighter in coloration, reddish brown; mesoscutellum brown; abdominal segments 3–5 reddish brown with dark brown areas medially and on paratergites; segment 6 brown except posterior 1/6 reddish brown; segment 7 brown on anterior half and reddish brown on posterior half; segment 8 reddish brown except anterior 1/3 brown. Head with 4–5 rows of large punctures on each side of impunctate middle epicranium; head with 4–5 rows of smaller punctures posteriorly; punctures contiguous; head appearing glossy due to lack of closely spaced micropunctures and microsculpture. Head transverse, width/length ratio = 1.25; antennomeres 1–6 longer than wide; antennomeres 7–8 subquadrate; antennomeres 9–10 slightly transverse. Neck with 4–5 irregular rows of small punctures. Pronotum longer than wide, width/length ratio = 0.83; with large, numerous, contiguous to virtually contiguous umbilicate punctures over most of pronotal surface; with impunctate midline, midline wider anteriorly than posteriorly; punctures round, not longitudinally elongate; punctures contiguous but not confluent; pronotum with polygon-shaped microsculpture on anterolateral corners. Pronotum weakly concave posteriorly. Pronotum/elytra length ratio = 0.96; elytra with large, dense punctures. Abdominal tergites with sparse, small punctures; sternites with denser larger punctures. Males with broad shallow emargination on sternite 7; small, round porose structure present positioned anteriorly on sternite 7; sternite 8 with broad shallow emargination; females with no secondary sexual structures. Aedeagus as in Figs. 6–7; paramere asymmetrical, in dorsal view converging to narrow rounded apex; in lateral view paramere becoming narrower medially to apex; paramere with two small apical teeth; paramere without peg setae; paramere shorter and narrower than median lobe; median lobe asymmetrical, in dorsal view, converging to narrow rounded tip; in lateral view median lobe becoming narrower medially, converging to upwards apex; medial lobe ventrally with to small teeth, almost beneath the parameral teeth. Distribution. Known from the department of Madre de Dios in Peru. Habitat. Philothalpus anapodos was collected in flight intercept traps in lowland tropical rainforest. Etymology. The specific epithet is derived from the Greek word ανάποδος (upside down) and refers to the shape of the median lobe.

Published as part of Chatzimanolis, Stylianos, 2020, Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae), pp. 129-138 in Zootaxa 4742 (1) on pages 130-131, DOI: 10.11646/zootaxa.4742.1.8, http://zenodo.org/record/3674539