Key to species of Ochodaeus in Madagascar 1. Pronotum everywhere strongly, densely granulate with elevated tubercles (Fig. 13)........ 2 — Pronotum either entirely smooth and punctate or with weak granulation confined to disc (Fig. 14)........................................................................ 4 2(1). Elytral tubercles on intervals sparse; body length less than 4 mm.... O. pygmaeus Paulian — Elytral tubercles dense; body length greater than 4 mm............................... 3 3(2). Median thickening of clypeal bead variable, lateral thickening always evident (Fig. 15); body usually> 9.0 mm....................................... O. isoanalensis Paulian — Median thickening of clypeal bead simply convex, bead thin laterally (Fig. 16); body length usually 4.5–8.0 mm................................................... O. miliaris Klug 4(1). Pronotal surface simply punctate............. O. modopunctatus Paulsen, new species — Pronotal surface weakly granulate on disc.......................................... 5 5(4). Clypeal bead evenly curved (Fig. 17). Mentum deeply, entirely furrowed (Fig. 5). Males usually with 2 pollices on protibia.................. O. polypollicatus Paulsen, new species — Clypeal bead wavy (Fig. 18). Mentum with furrow not reaching past middle (Fig. 6). Males with a single pollex on protibia...................... O. meandrus Paulsen, new species

Published as part of Paulsen, M. J., 2019, Three new Madagascan species of Ochodaeus Dejean (Coleoptera: Scarabaeoidea: Ochodaeidae), pp. 1-14 in Insecta Mundi 2019 (684) on page 11, DOI: 10.5281/zenodo.2579565