3.16.: Dichotomius (Cephagonus) gardneri new species (Figure 37 (i-l)) Diagnosis. separated from other in the group by the following combined characters: males pronotum central emargination on anterior portion; males cephalic carina inserted both eyesı transversal with acute apex (similar to D. aielloae and D. asenjoi); elytral striae and puncture deeply impressed; males paramera lacking basal and longitudinal excavations and paramera subgenital plate taking 1/5 of the ventral surface. ● HOLOTYPE (male): BRASIL: Pará. Monte Dourado. 0³60 ʹ 20 ʹ ’ Sı 52³59 ʹ 07 ” W. Fezes. iv- 2006. T Gardner & MIM Hernandéz [at CEMT]. ● PARATYPES: BRASIL. Pará. Óbidos. 1878. M Nathan [1 Ƌ at CEMT]; same municipality but xi-1989 [1 ♀ at CEMT]. Description (Holotypeı male). BL: 17 mmı PW: 9 mm. Colour: blackı shiny. Head: fronto-clypealı genal and inter-ocular surfaces smooth. Few weakly impressed fine punctures near eyes. clypeo-genal angle roundedı obtuse (100³). Cephalic carina longer than wideı conical with rounded apex. Pronotum: disc shinyı with central emargination – same width as interocular space. Posterior margin with a row of ocellate punctures. Metasternum: sides and margins of anterior lobe covered by long hair. Anterior portion of anterior lobe with ocellate setigerous punctures. Setigerous punctures equally spaced at sides. Elytra: striae deeply impressed having coarse punctures spaced by their diameter. Interstriae convexı disc lacking microsculpture with very fine punctures (20 x). Abdomen: ventrites with ocellate punctures concentrate at sides. Puncture of each ventrite interrupted medially as well as groove along the posterior margin of sixth ventrite. Aedeagus (Figure 37 (j-l)): dorsallyı paramere apexes roundedı convergent; longitudinal and basal excavations lacking; membranous part taking half of dorsal surface. Laterallyı subtle declivity from basis towards apex; lacking central excavation. Ventrallyı subgenital plate with rounded apexes and taking 1/5 of ventral surface. Morphological variation. male BL is 17 mmı PW: 9 – 10 mm. Female BL: 16 mmı PW: 11 mmı distinguished from males as follows: Head: fronto-clypeal surface striated. Surface posterior to cephalic carina smooth. Cephalic carina transversalı with straight apex. Pronotum: simply convexı lacking lobes. Elytra: striae weakly impressedı punctures separated by twice their diameter. Blue sheen present near basis. Abdomen: sixth ventrite bearing a pair of tubercles on central portion. Distribution. Brazilı state of Pará (Figure 34). Etymology. named after Toby Gardner (Stockholm Environment Institute)ı who collected the type series.

Published as part of Nunes, Rafael V. & Vaz-de-Mello, Fernando Z., 2019, Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini), pp. 2231-2351 in Journal of Natural History 53 (37) on pages 2322-2323, DOI: 10.1080/00222933.2019.1692088, http://zenodo.org/record/3665519