Cardiocondyla nuda (Mayr 1866) Tab. 2 Leptothorax nudus Mayr 1866: 508. Lectotype worker: Ovalau, Viti-Island (labels " Ovalau, Godeff. ", " 2768 ov " and " nuda G.Mayr, Type ") [NHMW, antweb.org images of specimen CASENT0919732] (examined). Combination in Cardiocondyla: Forel 1881: 6. All material examined. A total of 38 nest samples with 62 workers were subject to NUMOBAT investigation. Australia: New South Wales: Bulli, 1915.05.xx, [-34.31, 150.93]; Northern Territory: Gove, 1982.07.xx, [- 11.30, 132.30]; Queensland: Cairns: Esplanade Park, 2014.07.12, No AUS5 (GenBank LT718195), No AUS6 (GenBank LT718196), No AUS7 (GenBank LT718197), No AUS15 (GenBank LT718201), No AUS17 (GenBank LT718202), [-16.910, 145.767]; Queensland: Giru, 1981.xx.xx, [-19.51, 147.13]; Queensland: Kuranda, 1950.11.0 1, [-16.820, 145.630]; Fiji Islands: Ovalau, pre 1866.xx.xx (leg. Godeff), No 2768, type Cardiocondyla nuda, [-17.68,178.79]; Saiaro, pre 1960 (leg. W.M. Mann), [-17.90, 178.00]; Viti Levu, 1976.04.17, [-17.83, 177.94]. Micronesia: Pohnpei, 1994.07.30, [6.88, 158.23]; Pohnpei, 2000.03.19, [6.88, 158.23], Pohnpei, Ace, 1995.11.26, [6.97, 158.22]; Pohnpei, clear area, 1995.11.26, [6.97, 158.22]. New Hebrides (= Vanuatu): Aneityum, 1930.09.xx, [-16.50, 167.60]. Papua New Guinea: Haia- 10 km SSE, 2003.02.15, [-6.77, 145.02]; Huon: lower Busu River, [-6.73, 147.050]; Lae- 16 km NW, 1972.06.12, [-6.640, 146.89]; Manus, Ndrova Island, 1972.05.0 7, [-2.22, 147.24]; Morobe, Bulolo, 1967.12.18, [-7.76, 147.59]; Okapa- 15 km NW, Moife, 1959.10.xx, [-6.44, 145.51]; Samoa: Samoa Upolu, pre 1866.xx.xx (leg. Godeff), [-13.92, -171.74]; Tutuila: Pago-Pago, 1923.01.10, [-14.28, -170.70]; Upolu: Alagaogao, 1962.03.19, [-13.90, -171.70]; Upolu: Le Mafa, 1962.03.30, [- 13.88, -171.90]; Solomon Islands: Guadalcanal, 1964.xx.xx, [-9.64, 160.19]; Honiara, 1954.06.16, [-9.43, 159.94]; Isabel: Buala, 1964.12.14,, [-8.16, 159.62]; Malaita, 1965.xx.xx, [-9.05, 161.00]; Santa Catalina, 1933.07.xx, [-9.70, 160.00]. Tonga: Tonga Tapu, pre 1866.xx.xx (leg. Godeff), [-21.18, -175.22]. Wallis and Futuna: Futuna: Mt. Puke, 1965.xx.xx, [-14.276, -178.141]; Futuna: Nuku, 1965.03.xx, [-14.289, -178.166]; Nuku Hifala, 1965.03.30, [-14.30,-178.00]; Nuku Tapu, 1965.03/04, [-14.30, -178.00]; Uvea: Matu Utu, 1965.02.xx, [- 13.282, -176.177]. Geographic range. The distributional range of C. nuda extends over 6000 km in an east-west direction and includes the coast line of North and East Australia, Papua New Guinea and probably all Polynesian islands east to Samoa. The presence of populations in very isolated Polynesian islands (Fiji and Samoa) as early as the mid 19th century suggests that there was some natural way of dispersal over a huge Pacific area before modern ship traffic and goods transport developed. Diagnosis. see key. The clear cluster allocation of the lectotype is discussed in section 4.4. Biology. Three of six colonies excavated in 2014 in Cairns contained multiple fertile queens. Males are ergatoid, kill callow males and fight with adult males. Slightly larger, ergatoid males with stubby wings may occasionally occur (observations in the laboratory of J. Heinze).

Published as part of Seifert, Bernhard, Okita, Ichiro & Heinze, Jürgen, 2017, A taxonomic revision of the Cardiocondyla nuda group (Hymenoptera: Formicidae), pp. 324-356 in Zootaxa 4290 (2) on page 345, DOI: 10.11646/zootaxa.4290.2.4, http://zenodo.org/record/829077