Family Clupeidae Cuvier, 1816, new usage TAXA INCLUDED AND RELATIONSHIPS. — Clupea + Sprattus. UNAMBIGUOUS SYNAPOMORPHIES. — Participation of the pterotic in the sub-epioccipital fossa (58:1>0); anterior region of the mesethmoid keel perforated (62:0>1); olfactory nerve foramen of the lateral ethmoid surrounded by the lateral ethmoid and mesethmoid (66:1>2); teeth on the vomer present (69:0>1); ventral process of the quadrate short (128:0>1); foramen on the anterior ceratohyal produced by the overlap between lateral and medial foramina of the hyoid arterial duct present (140:0>1); foramen in the fourth epibranchial present (163:0>1); posterior concavity on the fourth epibranchial (166:0>1); holes on the surface of the coracoid present (272:0>1); keel of the pelvic scute absent (332:1>0); more than 30 abdominal scutes (341:3>4); focus of the body scales in postero-central position (360:2>3). SUPPORT. — Relative Bremer = 67%.

Published as part of Marramà, Giuseppe & Carnevale, Giorgio, 2026, A phylogeny and revised classification of clupeomorph fishes (Teleostei, Otomorpha), pp. 401-556 in Zoosystema 48 (17) on pages 521-522, DOI: 10.5252/zoosystema2026v48a17, http://zenodo.org/record/21479247