Piogaster variegata Bass, Bennett & Schwarzfeld sp. nov.Figs 20, 21Diagnosis.Piogaster variegata can be distinguished from its congeners by possession of a combination of the following: 1) mesoscutum matte and pustulate (Fig. 20 E); 2) HW 2 / Cu absent with no angulation in 1 / Cu & cu-a, only a very weak curve posterior to midheight present (Fig. 21 F); 3) T 1 – T 5 matte and granulate (Figs 21 G, 21 H); 4) metasomal T 2 – T 6 variegated: anterior 0.8 of each segment pale brown with dark brown spots, posterior 0.2 white (Fig. 21 H). Piogaster variegata is similar to other North American specimens (P. maculata and undescribed male Piogaster from British Columbia) as T 1 – T 5 of the metasoma are granulate and the abscissa 2 / Cu of the hind wing is absent. It can be distinguished by the variegated white and brown colour of the metasoma (Figs 21 G, 21 H) (tergites unicolorous with only narrow, posterior lighter bands in the other Nearctic specimens) (Figs 12 D, 23 C).Description.Adult. Female. Body length 4.3 mm. FW length 3.8 mm. Head. Antennae with 21 flagellomeres. Clypeus 2.0 × as wide as high, matte, granulate with sparse long white setae (Fig. 20 C). Face, frons and vertex matte, granulate with dense short white setae (Figs 20 C, 20 D). Occipital carina complete (Fig. 20 D). MSL 1.3 × as long as BWM. OOD 1.6 × as long as LOD. Mesosoma. Pronotum with epomia absent; matte, granulate, with dense medium length white setae (Fig. 21 A). Mesoscutum (Fig. 20 E), mesopleuron (Fig. 21 B), metapleuron, and propodeum (Fig. 21 D) matte, pustulate, with sparse to dense short to medium length white setae. Scutellum matte, granulate, with dense short white setae (Fig. 20 E). Propodeum with pleural carina complete; lateral longitudinal carina present in posterior 0.6, all other propodeal carina absent (Fig. 21 D). Wings. Fore wing. Vein Rs + M without ramellus extending into cell 1 M + 1 R 1 (Fig. 21 E). Vein 2 rs-m 0.6 × as long as M between 2 rs-m and 2 m-cu (Fig. 21 E). Vein 2 m-cu not thickened or angulate between bullae (Fig. 21 E). Hind wing. Vein 1 / Cu & cu-a slightly inclivous, slightly curved posteriorly, but not angulate, 2 / Cu absent (Fig. 21 F). Metasoma. T 1 matte, granulate with dense short white setae (Fig. 21 G). T 1 median dorsal carina present anteriorly but short, 0.1 × length of T 1; dorsolateral carina present anteriorly, 0.6 × length of T 1, absent posteriorly (Fig. 21 C). T 2 – T 5 matte, granulate with dense medium length white setae (Fig. 21 H). T 6 matte, granulate, with dense long white setae. T 7 matte, granulate and sparsely punctulate with dense long brown setae (Fig. 21 H). T 8 subpolished, punctulate with dense long brown setae. Tergites without grooves or tubercles (Fig. 21 H). Ovipositor sheath 0.6 × as long as hind tibia. Colour. Face white except stained with light brown near medial tubercle (Fig. 20 C). Clypeus dark brown basally, yellow apically (Fig. 20 C). Frons yellow-orange except white in orbits (Fig. 20 D). Vertex orange medially between lateral ocelli and posteriorly near occipital carina, these areas with a few small scattered light brown dots, sublaterally with white, subtriangular mark that is contiguous with white inner orbits of frons, laterally light brown with brown dots (Fig. 20 D). Gena white with dense, medium brown dots in dorsal half and posteriorly medium brown (Fig. 20 A). Occiput dark orange (Fig. 20 B). Mandible medium brown with transverse dark brown median band. Maxillary palps with segment 1 white, remaining segments light brown. Labial palps uniform white. Antenna light brown, scape, pedicel and flagellomeres 1–2 white anteriorly (Figs 20 A – C). Pronotum variegated white and medium brown. Tegula white. Mesoscutum predominantly orange, with a white H-shaped mark in the location where the notauli would be, the shape formed by sublateral, longitudinal white stripes joined by a transverse white mark just posterior to middle, lateral edges of mesoscutum narrowly white, posteromedially medium brown (Fig. 20 E). Scutellum white with anteromedial medium-brown spot (Fig. 20 E), axillary troughs orange-yellow in anterior half, white in posterior half (Fig. 20 E). Post-scutellum white, axillary troughs medium brown anteriorly, light brown posteriorly (Fig. 21 D). Mesopleuron variegated white and medium brown, ventrally light brown, area of sternaulus brown-orange. Metapleuron and propodeum (Fig. 21 D) variegated white and medium brown. Mesosternum light brown. Wings hyaline, veins including stigma white, Sc + R yellow-orange. Legs predominantly white. Fore and middle coxae with some small brown spots (two near base, one near apex), hind coxa with more extensive brown spots, laterally coalescing into a longitudinal band. Trochanters with small brown spots anteriorly and posteriorly that coalesce into longitudinal bands, trochantellus white with a small brown spot basally. Femora with small medium brown spots ventrally and laterally, dorsally coalescing into two longitudinal stripes anterodorsally and posterodorsally (Fig. 20 A). Fore tibia with a medium brown stripe on dorsal surface, and middle and hind tibiae with brown spots partly coalesced into longitudinal stripe dorsally. Tarsi medium brown dorsally, yellow-white ventrally. Metasomal tergites variegated white, yellow-white, and medium brown, most noticeably, round, slightly darker brown spots sublaterally near anterior margin of T 1–5 (Fig. 21 H). Ovipositor sheath white (Fig. 21 H).Male. Unknown.Distribution.Fig. 29. Canada (Alberta). This is the first record of Piogaster in both Canada and Alberta.Biology.Unknown.Material examined.Holotype. Canada • ♀; Alberta, EMEND site, 48 km NW of Dixonville; 56.7499°N, 118.3258°W; 29. vii – 12. viii. 2008; M. Schwarzfeld; Malaise trap, 853-1; deciduous forest, 50 % retention; UASM 327171; [CNC].Condition of type: Intact.Etymology.The epithet variegata is a descriptive name referring to the variegated colour patterns on the body, particularly the metasomal tergites, that are unique to this species in the genus Piogaster.Comments.The holotype of P. variegata is probably the most distinctive specimen of Piogaster known, with its predominantly variegated white and brown colouration and H-shaped white marking on the mesoscutum. These variegated spots are only known in some other specimens on the legs (as in P. maculata), not the body. With respect to distribution, this specimen and the unidentified male from British Columbia represent a huge range extension (2900 km distant between the type localities of P. maculata and P. variegata). This demonstrates the great range of the genus Piogaster within the Holarctic region and highlights the need for much more collecting of wasps and jumping spiders which will hopefully fill in gaps in the current species ranges and likely discover many more undescribed species.