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Phengodes (Phengodella) dichoclypeus Uchima-Taborda, Vega-Badillo & Zaragoza-Caballero, 2026, sp. nov.

Authors: Uchima-Taborda, Diego; Vega-Badillo, Viridiana; Zaragoza-Caballero, Santiago;

Phengodes (Phengodella) dichoclypeus Uchima-Taborda, Vega-Badillo & Zaragoza-Caballero, 2026, sp. nov.

Abstract

Phengodes (Phengodella) dichoclypeus sp. nov. Uchima-Taborda, Vega-Badillo & Zaragoza-Caballero(Figs: 11A–C, 18G–I, 24G–J, 27S–U)Description:Male. General Body: Body length 15, body width at shoulder height 3. Body color ochre yellow to honey yellow; except for around eyes, temples and apical half of mandibles brownish; apical third of elytron darkened, especially on borders; last three ventrites with brown areas. These ventrites with a dark brown triangular macula that may be divided on the penultimate ventrite; this macula is always present and divided (Fig 11A–C).Head (Fig 18G–I): Head 2.56 times wider (2.56) than long (1), feebly wider than pronotum (2.50). Eye 1.176 in length, 1.176 in height (lateral view), dorsal width 0.64 (dorsal view) and 0.76 in ventral width (ventral view); in lateral view 0.68 times the temple (1.73), eye posterior border concave and gena still visible, being one fifth of eye height (Fig 18I), in dorsal view interocular distance (2.058) is 3.216 times eye width, in ventral 1.4 times eye width. Antennal tubercles abrupt pronounced and close, covered by superficial punctures; interantennal distance (0.7) 1.59 times antennal fossae width (0.44), and lesser than scape length (0.625); antennal length (9.5) ending after elytral apex, and above the apex of third abdominal segment; scape widened in the middle and towards the apex, antennomeres I–III length (1.03), II conical (0,20), longer than III, latter cupuliform, compressed (0,17), ventrally projected, IV–VI (1.50), IV<V<VI, flabellae of these antennomeres 6–7 times longer than respective antennomere, antennomere XII (3.72) 3.98 times longer than XI (0.93).Vertex with longitudinal carinae converging to center, without a central longitudinal sulcus. Longitudinal sulcus in the frons absent; fronto-clypeal suture straight; clypeus apically notched, divided, briefly fused near base, where a middle line is present; clypeal lobes fusiform; clypeal puncture diameter wider than lumen, more punctate in external half, with abundant setae. Mandibles long (1.53), 1.53 times head length, thin and dorsoventrally rounded; mandible widen at base, then thinning from curvature toward apex, setae and punctures present up to apex, without marks on external or internal borders. Galea developed, lanceolated, with 5–6 setae at apex; maxillary palpi (2.36) long and thin, 2.03 times labial palpi length (1.16), maxillary palpi formula I<II>III<IV, II>IV, palpomere II (0.70) 3.5 times I (0.20); labial palpi formula 0<I<II<III, palpomere I (0.23) 1.4 times palpiger (0.17) (Fig 18H).Thorax: Pronotum transverse, 1.51 times wider at base (2.5) than long (1.66), wider at middle (2.66); anterior and lateral borders rounded, posterior border undulated and discontinuous, with a carinae that occupies extremes and center of the discontinuity, posterior corners right-angled and angled. Lateral expansion wide (0.33), pronotum width 7.75 times expansion width, expansions widened near apex, narrow near posterior border. Pronotal disc transverse, without longitudinal sulcus, tubercles absent, punctures superficial and dense, whose lumen is less than its diameter, lumen smooth. Scutellum 2.91 times longer (0.67) than wide (0.23). Elytra apically expanded, 2.5 times longer (3.5) than wide (1.4) and 2.1 times longer than pronotum, ending at the apex of the second abdominal segment; elytral disc with two visible costae, punctuation dense at apex (Fig 18G), membranous wings 2.07 times longer (10.33) than wide (5.0). Tibial apex with uniform spines, tarsal arolium elongated, present only in fourth tarsomere, pulvilli formed by two setae, tarsal claw glabrous.Abdomen: Abdominal segments weakly expanded and covered by minute yellow hairs; posterior border of tergite VIII rounded, X straight; posterior border of ventrite VII undulated, without medial projection, VIII emarginate (Fig 24G–J). Aedeagus long (1.56), symmetric; central lobe longer than parameres. Parameres 3.5 times longer (1.16) than wider (0.33); apex with two pronounced teeth, dorsal absent, central and ventral equal in size, internal border of paramere not sinuate, flagellum (6.7) 4.29 times aedeagus length (Fig 27S–U).Female: Unknown.Diagnosis: Galea lanceolated and acute with few hairs at apex, clypeus divided and clypeal lobes fusiform, mandibles thin and dorsoventrally rounded, antennae feebly surpassing elytra, ventrite VII without projection, paramere with two apical denticles, central wider than ventral, additionally the abdominal color pattern is stable.Remarks: This species is similar to P. (Pla.) huatulcoensis sp. nov. from which it differs by the following characteristics: in P. (Pla.) diploclypeus, the clypeus is divided and apically notched, pronotum is wider and the paramere base is ventrally straight while the anterior border of the paramere is angled. In contrast, in P. (Pla.) huatulcoensis, the clypeus is entire, not divided and not apically notched, the pronotum is narrower, the paramere base is dorsally widened and the anterior end of the paramere rounded. The coloration pattern is also useful to separate them since P. (Pla.) huatulcoensis has a different color pattern on the abdomen. P. (Pla.) diploclypeus is similar to P. (Pla.) succinacius but it differs mainly by the genitalia, which is more rounded and stouter in P. (Pla.) succinacius which also has the apical denticles closer and pointier than in P. (Pla.) diploclypeus. It is also similar to P. (Ph.) chamelensis but can be differentiated by the pronotum and head form, and the galea that is lanceolated in P. (Pla.) diploclypeus.Variation: Size 10.09–15, width at shoulder high 2.4–3, head width 2–2.56; interantennal distance 0.55–0.73, interocular distance 1.29–1.53; pronotum length 1.29–1.66, pronotum width 1.8–2.63, elytra length 3.14–3.5, and elytra width 1.11–1.43. Clypeus usually completely divided into two halves, if fused, then just briefly at base and middle line still evident. When observed in liquid this middle line is translucent and membranous, making the separation more evident. In dry specimens the clypeus is concave.Distribution and ecology: Oaxaca (Fig 30). Encountered in Huatulco in tropical dry forest habitat, this species shows activity from May to July, which are the beginning of the rainy season reported for the area (Noguera et al. 2018). This pattern could suggest a phenology coordinated with the start of the rainy season. However, more studies are needed to confirm this.Etymology: dicho - Gr. in two, by half, the name P. dichoclypeus refers to the divided clypeus.Examined material: HOLOTYPE: MEXICO: Oaxaca, Huatulco 4,6 KM NE Entronq Carr Salina Cruz/ Santa Cruz, 08–13/05/2005, 15°48’43.3”N, 96°10’19,7 O, 194 masl Trampa Malaise, S. Zaragoza, F. Noguera, E. Gonzalez, E. Ramírez, (CNIN) (1 ♂). PARATYPES: MEXICO: Oaxaca, Huatulco, 4, 5 km NE Estronq Carr Salina Cruz/Santa Cruz, 08–13/05/2005, 15°48’40,7”N, 96°10’18,0”O, 187 masl, Trampa Malaise, S. Zaragoza, F. Noguera, E. Gonzalez, E. Ramírez, (CNIN) (2 ♂); idem except for: Huatulco 4,6 KM NE Entronq Carr Salina Cruz/ Santa Cruz, 08–13/05/2005, 15°48’43.3”N, 96°10’19,7 O, 194 masl (2 ♂); idem except for: Huatulco, 5,1 KM NE Entronq Carr Salina Cruz/Santa Cruz, 15°48’54,1”N, 96°10’28,6”O, 179 masl, (1 ♂); idem except for: Huatulco, El Carnero, 3,7 KM NE Entronq Carr Salina Cruz/Santa Cruz, 08–13/05/2005, 15°48’21,4”N, 96°10’04,0”O, 90 masl (1 ♂); idem except for: 06–11/06/2005, 15°48’21,4”N, 96°10’04,0”O, 90 masl, (2 ♂); idem except for: Oaxaca, Huatulco Entronque Carr Salina Cruz/Santa Cruz, 06–11/07/2005, 15°48’33,3”N, 96°10’11,7”O, 204 masl (3 ♂)

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