Parastenomyia waynei Kameneva and V. Korneyev, new species (Figs. 1, 5, 9–36) urn:lsid:zoobank.org:pub: EF129D02-A6E4- 4AB0-82FB-952D8F0C9145 Diagnosis.—This species can be readily recognized by the key characters: the wing pattern with the preapical hyaline interspace parallel-sided, uniformly narrow, and anteriorly extended into cell r 1, in combination with the relatively narrow prediscal hyaline interspace and two unequal midtibial setae. It is similar in wing pattern to P. willii, from which it can be easily recognized by having only two shorter midtibial setae as in Fig. 52 (vs. 3–4 very long setae as in Figs. 53–54 in P. willii). Description.—Medium-sized, body length 3.5–5.5 mm, subshining black flies with banded wings. Head (Figs. 9–12, 15–17): length: height: width = 1: 1.45: 2.0 (1: 1.3: 1.7 in paratype ♂); mostly brownish-yellow, with black ocellar triangle, vertical plates and most of occiput (usually with brownish-yellow triangle ventral to postocellar setae); frons 0.95 × as long as wide (1.0 in paratype ♂), with 3–6 irregular frontal setae, posterior inclinate frontal seta often between vertical plates and ocellar triangle. Eye very large, 1.4–1.5 × as high as long (1.3 × in paratype ♂) and 6.0–6.5 × as high as gena (4.5 × in paratype ♂). Epistome yellow-brown to dark brown. Postpedicel 1.5 × as long as wide. Clypeus black, half as high as epistome, entirely white microtrichose. Palpus brownish-yellow, long and densely setulose, brown and white microtrichose. Thorax (Figs. 9, 12, 16, 19): Mostly black or dark brown, sometimes with yellowish-brown postpronotum and notopleural triangle. Scutum sparsely microtrichose, shining black with bluish sheen from posterior view and matt from anterodorsal view; pleura with somewhat denser microtrichia; scutellum flat, dark brown to black, densely transversely shagreened, subshining, very sparsely microtrichose dorsally, denser microtrichose on sides. Legs: mostly black to brown; fore coxa brownish-yellow to brown, all coxae densely microtrichose. Wing (Figs. 1, 5, 13, 14): ♀ 3.0–4.0 mm (mean = 3.7 mm); holotype ♀ 3.6; ♂ 4.0 mm long, 2.5–2.7 times as long as wide. Vein R 1 in apical section with 5 (♂ — Fig. 18) to 7–10 (♀) setulae dorsally along pterostigma. Basicostal cell hyaline; subbasal crossband complete, reaching from humeral crossvein and basal 1/6 of costal cell to alula; alula pale grey microtrichose; prediscal hyaline interspace discal crossband from apical 1/6 of costal cell and whole pterostigma slightly widening to posterior margin. Medial hyaline interspace almost completely white microtrichose (with grayish microtrichia at posterior margin), 0.5–1 × as wide as discal crossband and 0.6–2 × as wide as subapical band (2.5 × in ♂). Subapical crossband as wide as vein dm-m length, narrowly connected to apical band in cell r 1; preapical hyaline interspace narrow,0.5–1×as wide as subapical dark band, parallel-sided throughout its length, anteriorly reaching only middle of cell r 1 width, and separated from posterior margin in cell m 1 by gray microtrichose area. Apical band very wide, D-shaped, with proximal margin perpendicular to veins R 4+5 and M 1. Male genitalia as in Figs. 21–27 and described for the genus. Female genitalia as in Figs. 28–36. Aculeus 6 × as long as wide, its apex (Figs. 11, 35) very narrowly pointed: distance from basal most subapical setulae to apex 3 × longer than distance between these setulae. Distribution.— Costa Rica. Type data.— Holotype female, COSTA RICA: Limón: Bribri (4 km E, 50 m, 10º00'N, 83º15'W, Malaise trap, Jul–Sep 1990, P. Hanson (LACM). Paratypes: COSTA RICA: Limón: Valle de la Estrella Reserva Biol., Hitoy Cerere, 140 m, 17 Jul–17 Jun 1999, F. Umafia, 1f (Malaise trap, L_N_184600_643 400 #52861) (INB 0003307747 INBIO CRI COSTA RICA) (MNCR); Cartago: Madreselva, Fincos los Lagos, A.C., Amistad, 2600 m. Dec 2000, M.M. Chavarría, 1f (Malaise trap, L_N_184450_550050 #2602) (INB 001864921 INBIO CRI COSTA RICA) (MNCR); Turrialba, 15–19 Jul 1965, P. J. Spangler, 1m (LACM); Alajuela: RNVS Cafío Negro, 20 m, 13-20 Feb 1994, K.L. Martinez, 1f (Malaise trap, L_N_319100_450100 #2887) (INB 003033834 INBIO CRI COSTA RICA) (MNKB) [dissected]; Puntarenas: Golfito, Pque. Nal. Corcovando, Argujos, Charcos, 600m, 17 Apr–17 May 1999, J. Azofeifa, 1?, 2f (Malaise trap, L_N_276350_523500 #52766, 52776, 52781) (INB 003307520, 003307525, 003307538, INBIO CRI COSTA RICA) (MNCR); Golfito, Pque. Nal., Est. Argujos, Cerra Rincón, 745m, 15 Feb 2000, J. Azofeifa, 1f (Malaise trap, L_N_275500_522000 #55202) (INB 003308237, INBIO CRI COSTA RICA) (MNCR); P.N. Manuel Antonio, 80 m, Quepos, Dec 1991, G. Varela, 1m [dissected] (L_S_370900_448800 (INB 000528637, INBIO CRI COSTA RICA) (SIZK); Heredia: Estatio La Selva, 15 Jul 1993, ALAS, Malaise trap M/09/159, 1f (CSCA); idem, 14 Aug 1993, ALAS, Malaise trap M/01/180, 1f (CSCA).

Published as part of Kameneva, Elena P., Korneyev, Severyn V. & Korneyev, Valery A., 2025, Parastenomyia New Genus (Diptera: Ulidiidae: Lipsanini) From Costa Rica, Ecuador, And Peru, pp. 485-500 in Proceedings of the Entomological Society of Washington 127 (3) on page 496, DOI: 10.4289/0013-8797.127.3.485, http://zenodo.org/record/18484343