GenusCrassignathaWunderlich, 1995 Crassignatha Wunderlich, 1995: 546; Miller et al. 2009: 68; Li et al. 2020: 66. Type species. Crassignatha haeneli Wunderlich, 1995, by original designation, from Malaysia. Diagnosis and description. See Li et al. 2020: 66–67 in detail. Composition. Crassignatha baihua Lin & Li, 2020 (♂ ♀), C. bangbie Lin & Li, 2020 (♂ ♀), C. bicorniventris (Lin & Li, 2009) (♀), C. bispina (Lin, Pham & Li, 2009) (♂ ♀), C. changyan Lin & Li, 2020 (♀), C. danaugirangensis Miller et al., 2014 (♂ ♀), C. dongnai Lin & Li, 2020 (♀), C. ertou Miller, Griswold & Yin, 2009 (♂ ♀), C. gucheng Lin & Li, 2020 (♂ ♀), C. gudu Miller, Griswold & Yin, 2009 (♀), C. haeneli Wunderlich, 1995 (♂), C. hekou Wu & Lin, sp. nov. (♂ ♀), C. kishidai (Shinkai, 2009) (♂ ♀), C. liangdu Wu & Lin, sp. nov. (♂ ♀), C. mengla Lin & Li, 2020 (♂ ♀), C. nantou Lin & Li, 2020 (♂ ♀), C. nasalis Lin & Li, 2020 (♂ ♀), C. panlong Wu & Lin, sp. nov. (♂ ♀), C. pianma Miller, Griswold & Yin, 2009 (♂ ♀), C. qingxu Wu & Lin, sp. nov. (♂ ♀), C. quadriventris (Lin & Li, 2009) (♂ ♀), C. quanqu Miller, Griswold & Yin, 2009 (♂ ♀), C. rostriformis Lin & Li, 2020 (♂ ♀), C. seedam Rivera-Quiroz, Petcharad & Miller, 2021 (♀), C. seeliam Rivera-Quiroz, Petcharad & Miller, 2021 (♂ ♀), C. shiluensis (Lin & Li, 2009) (♂ ♀), C. shunani Lin & Li, 2020 (♂ ♀), C. si Lin & Li, 2020 (♂ ♀), C. thamphra Lin & Li, 2020 (♀), C. xichou Lin & Li, 2020 (♀), C. yamu Miller, Griswold & Yin, 2009 (♂ ♀), and C. yinzhi Miller, Griswold & Yin, 2009 (♂ ♀). Distribution. Southern China (Guizhou, Yunnan, Hainan, and Taiwan), Central Japan (Honshu, Shikoku), Vietnam, Thailand, and Malaysia. Comments. The spider genus Crassignatha was initially placed in Synaphridae (Wunderlich 1995), later cataloged within Mysmenidae (WSC 2025), and subsequently transferred to Symphytognathidae, where it is currently classified (Miller et al. 2009; Lopardo and Hormiga 2015; Li et al. 2020). The genus now comprises 32 species, including the four newly described in this study, and is primarily distributed across the Far East and Southeast Asia. With this updated species count, Crassignatha surpasses both Symphytognatha Hickman, 1931, and Patu Marples, 1951, in species richness, making it the second-largest genus in Symphytognathidae. It is currently exceeded only by Anapistula Gertsch, 1941, which contains 34 recorded species (Brescovit and Cizauskas 2025). Furthermore, Crassignatha represents the most species-rich genus of Symphytognathidae known from China to date.

Published as part of Wu, Haihong & Lin, Yucheng, 2026, Phylogenetic classification of poorly known Crassignatha spiders (Araneae, Symphytognathidae), with descriptions of four new species, pp. 159-179 in Zoosystematics and Evolution 102 (1) on pages 159-179, DOI: 10.3897/zse.102.165834