Cycloryx granum (Godwin-Austen, 1893) Fig. 22F–O Alycaeus otiphorus var. Godwin-Austen, 1871: 93, pl. 5 fig. 6. Alycaeus (Dioryx) granum Godwin-Austen, 1893: 593–594. Alycaeus (Cycloryx) multicostulatus Godwin-Austen, 1914: 404, pl. 147 fig. 7. Syn. nov. Alycaeus (Cycloryx) burrailensis Godwin-Austen, 1914: 403 (in part). Alycaeus (Dioryx) granum – Godwin-Austen 1897: 4–5, pl. 63 fig. 6. Alycaeus (Alycaeus) granum – Kobelt 1902: 345. Alycaeus (Cycloryx) granum – Godwin-Austen 1914: 364. — Gude 1921: 279–280. Alycaeus (Cycloryx) multicostulatus – Gude 1921: 282–283. Cycloryx granum – Ramakrishna et al. 2010: 71. Cycloryx multicostulatus – Ramakrishna et al. 2010: 73. Pincerna granum – Páll-Gergely et al. 2020: 177. Pincerna multicostulata – Páll-Gergely et al. 2020: 181. Diagnosis Shell globular, ribs dense on the entire shell. Type material examined INDIA • 1 syntype of A. granum (Fig. 22F–J); Margarita, near Sadia, Assam; Godwin-Austen coll.; NHMUK 1903.7.1.2511 • 1 syntype of A. multicostulatus (Fig. 22K–O); Munipur, head of Lanier Rr; Godwin-Austen leg.; NHMUK 1903.7.1.2557. Type localities “Margarita, foot of Eastern Naga Hills” (A. granum); “Head of the Lanier River, Lahupa Naga Hills, N.E. Manipur ” (A. multicostulatus). Remarks Types of C. granum are much smaller than C. multicostulatus, but there is no difference between those shells other than shell size. Specimens from the Japvo Peak, Naga Hills (labelled as “ A. burrailensis var. minor ”, see also A. burrailensis) are intermediate in size between C. granum and C. multicostulatus. Therefore, we treat the latter as a junior synonym of the former.

Published as part of Páll-Gergely, Barna, Gojšina, Vukašin & Aravind, Neelavar Ananthram, 2025, A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae), pp. 1-158 in European Journal of Taxonomy 1029 on pages 40-41, DOI: 10.5852/ejt.2025.1029.3131, http://zenodo.org/record/17864673