54 b.Drymaria villosa subsp. palustris(Schltdl. & Cham.) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 227. 1961. ≡ Drymaria palustris Schltdl. & Cham., Linnaea 5: 232. 1830. Type: Mexico. Veracruz: In paludosis prope Jalapam, s. d., C. J. W. Schiede & F. Deppe 504 (lectotype, designated by Duke 1961, pg. 227: LE [LE 00001733 image!]; isolectotype LE [LE 00001732 image!]). ≡ Drymaria cordata var. palustris (Schltdl. & Cham.) Rohrb., Fl. Bras. 14 (2): 260. 1872. = Drymaria pauciflora Bartl. in Presl, Reliq. Haenk. 2 (1): 8. 1831. Type: Peru: in Peruviae montanis huanoccensibus, 1822, T. Haenke s. n. (lectotype, designated here: GOET [GOET 000582 image!]; isolectotype: PR [24306 image!]). = Drymaria townsendii B. L. Rob., Bot. Gaz. 30 (1): 58. 1900. Type: Mexico. Chihuahua: Sierra Madre, 5 miles southeast of Colonia Garcia, 7400 ft [alt. 2255.5 m], 5 August 1899, C. H. T. Townsend & C. M. Barber 231 (holotype: GH [00037745 image!]; isotypes: E [E 00326057 image!], F [102975 / F 0053306 F image!], K [K 000471738 image!], MO [3578839 / 144037 image!], NDG [16755 image!], NY [00342510 image!], US [735652/00103368 image!], US [00931459 image!]). ≡ Drymaria villosa f. townsendii (B. L. Rob.) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 228. 1961. = Drymaria nummularia Briq., Annuaire Conserv. Jard. Bot. Genève 13: 371. 1911. Type: Mexico. [Michoacán]: cordillere d’ Ario, 4000, Jun. – Oct. 1840, H. G. Galeotti 4416 (holotype: G [G 00226781 image!]; isotype: K [3862 / K 000471748 image!]). = Drymaria subsessilis M. E. Jones, Contr. W. Bot. 15: 125. 1929. Type: Mexico. Nayarit: in rivulets at Ixtlan, 19 February 1927, M. E. Jones 22848 (holotype: POM [162843/0002391 image!]; isotype: F [705822 / F 0053303 F image!]). = Drymaria sphagnophila Baehni & J. F. Macbr., Publ. Field Mus. Nat. Hist., Bot. Ser. 13 (2 / 2): 625. 1937. Type: Peru. Huánuco: Mito, 9000 ft [alt. 2743.2 m], 8–22 July 1922, J. F. MacBride 1542 (holotype F n. v.; isotype: US [1779692 / 00103362 image!]). = Drymaria villosa subsp. palustris var. perennis J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 228. 1961. Type: México. San Luis Potosí: Álvarez, s. d., E. Palmer 187 (holotype: MO n. v.). Distribution. Mexico, Guatemala, Honduras, El Salvador, Costa Rica, Panama, Venezuela, Colombia, Ecuador, Peru, Bolivia. Notes. Most of the type specimen deposited at B were destroyed in World War II (Hiepko 1987), and Duke (1961) mentioned that the holotype of Drymaria palustris at B was probably destroyed and designated as the lectotype the specimen cited above. This sheet bears an original label written by Chamisso stating “ Drymaria palustris n. sp. ” (handwriting confirmed by Denise Marx, HAL). However, the number “405” does not correspond to the number given in the protologue, which is “404.” Despite this discrepancy, apparently due to a labeling error, we believe that this specimen is original material and thus agree with Duke’s lectotypification. Although there are two specimens at LE, only one was labeled by Duke ([00001733]). We treat the other ([00001732]) as an isolectotype. For the same reason as mentioned for Drymaria apetala, we designate the specimen in GOET as the lectotype of Drymaria pauciflora.

Published as part of Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas & Montesinos-Tubée, Daniel B., 2025, A nomenclator of Drymaria (Caryophyllaceae), pp. 139-175 in PhytoKeys 266 on pages 139-175, DOI: 10.3897/phytokeys.266.162517