Raddaus tuberculatus (Rathbun, 1897) (Figs 6 F-H; 8; 16) Pseudothelphusa tuberculata Rathbun, 1897: 61; 1898: 511 (in key), 513, 535, and 537 (in list); 1904: 242 (in list); 1905: 299, pl. 13, fig. 9; 1906: 502 (in list). — Young 1900: 217 — Coifmann 1939: 109 (in list). Pseudothelphusa (Anaphyrmus) tuberculata – Smalley 1965: 651 (in list). Potamocarcinus (Raddaus) tuberculata – Pretzmann 1965: 3 (in list). Pseudothelphusa (Raddaus) tuberculatus – Pretzmann 1971: 20 (in list); 1972: 80, figs 461-464, 519-521. Raddaus tuberculatus – Rodríguez 1982: 103, figs 64i, 66a-e. — Villalobos F. 1982: 220 (in list). — Villalobos Hiriart & Álvarez 2008: 297 (in list). — Ng et al. 2008: 177 (in list). — Wehrtmann et al. 2016: 781 (in list). — Acevedo-Alonso & Cumberlidge 2022: 583 (Appendix 2, in list). TYPE MATERIAL EXAMINED. — Lectotype (designated by Smalley 1965). Guatemala • 1 ♂ (50.0:30.9, in bad condition); Suchitepéquez, stream of Saint Augustín, near Atitlán [? Finca San Agustím; 14°28’23.93”N, 91°12’53.26”W, 609 m elev.], Pacific drainage; date unknown, M. Bocourt; USNM 20047. Paralectotype. Guatemala • 1 ♀, Suchitepéquez, stream of Saint Augustín, near Atitlán [? Finca San Agustin; 14°28’23.93”N, 91°12’53.26”W, 609 m elev.], Pacific drainage; M. Bocourt leg.; MNHN Entry 844-1866; MNHN-IU-2017-8048 (= MNHNB5322). OTHER MATERIAL EXAMINED. — Guatemala • 1 ♂ (69.5:41.4); San Marcos, Finca El Porvenir; [14°57’43”N, 91°56’26”W]; 1370 m elev.; 22.III.1952; W. Reeder leg.; USNM 1089233 • 2 ♂ (18.9:13.5-35.1:22.5) 3♀ (22.6:14.9-35.7:22.9), San Marcos, sítio San Rafael, San Marcos; 14°56’21”N, 91°54’38”W; 991 m elev.; 7.VIII.2015, I. S. Wehrtmann & M. Orozco leg. UCR-MZ uncat. • 1♂ (23.5:17.7); Suchitepéquez, carretera a Chicacao; 14°27’50”N, 91°09’01”W; [471 m elev.]; 19.I.2014, I. S. Wehrtmann, M. Orozco, C. Magalhães & M. W. Dix leg. UCR-MZ uncat. DISTRIBUTION. — Guatemala (San Marcos, Suchitepéquez) (Fig. 16) and Mexico (Rathbun 1897, 1905; Pretzmann 1972; this study). DESCRIPTION OF G1 (Fig. 6 F-H) Somewhat slender, slightly arched in cephalic direction in mesial view, moderately compressed mesolaterally; caudal margin with median constriction in mesial view; cephalic margin gently concave in proximal 2/3, with few short setae subdistally; lateral surface with irregular rows of short to large setae in first half and few minute setae subdistally. Marginal plate fused subdistally to caudal surface. Marginal suture on mesial surface straight up to apex, with row of large to short marginal setae along proximal portion. Marginal process subrectangular, nearly as long as mesial margin of apex. Lateral suture incomplete, marked by distinct sulcus on caudal surface. Mesial process broad, unilobed in mesial view, subtriangular, narrower than width of main stem in mesial view, directed cephalad; proximal corner rounded, distal corner with small sharp spine pointing lateralad in lateral and apical-cephalic views; cephalic surface of mesial process distinctly concave in lateral and apical views. Apex oblong, situated apically, open, mesial and lateral margins about same height; field of apical spines moderately developed, with short small spines; spermatic channel opening on apex cavity, situated medianly. Cephalic surface with two strong, triangular, subdistal spines, pointing cephalad. REMARKS Rathbun (1897) described the species based on a male and a female at this time deposited in the holdings of the MNHN (Paris); the male specimen was later sent to the USNM collection (Washington, D.C.). When Smalley (1965) introduced the subgenus Pseudothelphusa (Anaphyrmos), he designated Raddaus tuberculatus as the type species because it was the first species he included in the subgenus to have been described from a male and, to consolidate his decision, he designated the specimen deposited in the USNM (20047) as lectotype. This specimen is in poor condition, completely disarticulated, the posterior ventral portion severely damaged, the posterior portion of the sternopleonal cavity broken, with both gonopods missing (they were not found in a small vial kept within the jar as usual) (Fig. 8).

Published as part of Magalhães, Célio & Wehrtmann, Ingo S., 2025, Taxonomic revision and distribution of the freshwater crabs of the family Pseudothelphusidae (Decapoda, Brachyura) from Guatemala, with the descriptions of two new species and keys to the species of all genera, pp. 581-615 in Zoosystema 47 (26) on page 601, DOI: 10.5252/zoosystema2025v47a26, http://zenodo.org/record/17432059