Zodarion modestum (Simon) (Figures 17, 24G–I, 25N, 26N, 27K) Enyo modesta Simon, 1870: 145 (♂). Zodarion modestum – Denis 1937: 32, pl. 6, figs 47–48 (♂). – Bosmans 1994: 136, figs 79– 80, 131–132 (♂ ♀). Examined type material Lectotype (♂) and paralectotype (♂): UNITED KINGDOM (IBERIA): Gibraltar; MNHN-AR1519, MNHN-AR1520. Other material examined SPAIN: 1 ♂; Zahara de los Atunes, Cádiz; 10 m a.s.l.; 9 April 1974; J. Murphy and F. Murphy leg.; wooded hillside; CJFM-677, G7572.686. 1 ♀; idem; 0 m a.s.l.; 14 April 1974; J. Murphy and F. Murphy leg.; sand dunes; CJFM-677, G7572.686. 2 ♀, 1 juv.; labelled as ‘ Málaga, Maro’; 100 m a.s.l.; May – April 1987; J. Murphy and F. Murphy leg.; stones, litter; CJFM-14343, G7572.14989. 1 ♂, 2 ♀; labelled as ‘Málaga, Frigiliana’; 20 m a.s.l.; 20 April 1987; J. Murphy and F. Murphy leg.; stones, litter; CJFM-14558, G7572.12808. 1 ♀; labelled as ‘Granada, Cádiz’; MNHN-AR1522. 3 ♂, 5 juv.; Rio Maitena, Granada; 28 May 2012; S. Pekár and S. Korenko leg.; CMU. 2 ♀, 1 juv.; Facinas, Cádiz; 36.145°N, 5.702°W; 2 December 2021; D. Ortiz and A. Avila leg.; urban grassland; CMU. 1 ♂; around pond of Western outskirt of Mont-Ras, Girona; 41.915°N, 3.141°W; 23 December 2021; D. Ortiz leg.; CMU. 4 ♂, 2 ♀; Casares, Málaga; 36.446°N, 5.279°W; 2 May 2024, S. Pekár and S. Shafaie leg.; CMU. Diagnosis Males of Z. modestum can be distinguished from the closely related Z. isabellinum by the absence of an embolar base (vs present; compare Figure 24H and Figure 24B) and the gradually narrowing embolus (vs a more slender embolus; compare Figure 24H and Figure 24B), while females can be distinguished by a sub-trapezoidal epigynal plate with a longitudinal, midline suture (vs epigynal plate sub-rectangular, with a longitudinal, concave median septum; compare Figures 25N, 27K and Figures 25L, 27I). Description For a detailed description of the habitus and male palp, refer to Bosmans (1994). Epigyne. (Figure 27K) Sub-trapezoidal epigynal plate, with a longitudinal, midline suture posteriorly. Epigynal plate edges sinuous in shape. Molecular data COI mitochondrial barcodes (GenBank accession): PQ119991, PQ119992. Maximum intraspecific COI divergence (2): 6.4%. Minimum interspecific divergence (to Z. isabellinum): 6.3%. Consensus COI distance to the closest species (Z. isabellinum): 3.4%. Distribution Spain, United Kingdom (Gibraltar).

Published as part of Shafaie, Sepideh, Pekár, Stano & Ortiz, David, 2025, Zodarion spiders (Araneae: Zodariidae): morphology, molecular characterisation and reassessment of the elegans, italicum, and pusio groups from Iberia and the Canary Islands, pp. 1825-1875 in Journal of Natural History 59 (25 - 28) on pages 1851-1857, DOI: 10.1080/00222933.2025.2498047, http://zenodo.org/record/17006654