Onitis feae Felsche, 1907 Figs 10A, 11E–H, 18 Onitis feae Felsche, 1907: 293 (original description). Onitis feae – Boucomont & Gillet 1921: 17, 18 (distribution). — Arrow 1931: 397 (key, description). — Janssens 1937: 75 (revision). — Paulian 1945: 141 (revision). — Singh et al. 2010: 1395 (list). — Gupta et al. 2015: 1037 (checklist). — Bezděk 2016: 179 (distribution). — Schoolmeesters 2022 (online catalogue). Differential diagnosis Onitis feae is distinct in having an elliptical clypeus and a sharply elevated frontal carina which is complete. Type material examined Cotype MYANMAR • ♂; “Asciuii Cheba, 1200-1300 m, L. Fea. 1–88; Co-Typus (red label); Coll. C. Felsche, Kauf 20, 1918; Staatl. Museum für Tierkunde Dresden; SNSD ”. Additional material examined (111 specimens) INDIA • ♂; Meghalaya, Shillong, village near Northeast Hill University (NEHU); 31 Oct. 2012; Seena Narayanan Karimbumkara leg.; AIMB / Co/Sc1000185 • 2 ♂♂, ♀; Arunachal Pradesh, Mobu; 27°14ʹ17.7324ʺ N, 93°32ʹ14.28ʺ E; elev. 876 m a.s.l.; 19 Oct. 2018; hand-picked, mithun dung; Priyadarsanan Dharma Rajan and team leg.; AIMB / Co/Sc1000186 to 1000188 • 6 ♂♂, 5 ♀♀; Arunachal Pradesh, Aalong; 28°8ʹ36.384ʺ N, 94°49ʹ33.34ʺ E; elev. 317 m a.s.l.; 14 Nov. 2019; hand-picked, mithun dung; Sneha Haridas and team leg.; AIMB / Co/Sc1000189-1000199 • 13 ♂♂, 12 ♀♀; Arunachal Pradesh, West Siang, Aalong forest; 28°8ʹ33.882ʺ N, 94°49ʹ36.1848ʺ E; elev. 359 m a.s.l.; 14 Nov. 2019; cow dung baited pitfall trap; Sneha Haridas and team leg.; AIMB / Co/Sc1000200 to 1000224 • 17 ♂♂, 22 ♀♀; Arunachal Pradesh, West Siang, Kaying forest; 28°24ʹ35.8668ʺ N, 94°41ʹ51.4896ʺ E; elev. 376 m a.s.l.; 15 Nov. 2019; cow dung baited pitfall trap; Sneha Haridas and team leg.; AIMB / Co/ Sc1000225 to 1000263 • 10 ♂♂, 15 ♀♀; Arunachal Pradesh, West Siang, Darak / Larmuk Pottom forest; 28°12ʹ17.568ʺ N, 94°35ʹ49.3008ʺ E; elev. 489 m a.s.l.; 18 Nov. 2019; mithun dung baited pitfall trap; Sneha Haridas and team leg.; AIMB / Co/Sc1000264 to 1000288 • 2 ♂♂, 2 ♀♀; Arunachal Pradesh, Boru Raksap / Yardi Rabe Supse Wildlife Sanctuary; 28°15ʹ56.1132ʺ N, 94°34ʹ58.008ʺ E; elev. 529 m a.s.l.; 24 Nov. 2019; mithun dung baited pitfall trap; Sneha Haridas and team leg.; AIMB / Co/Sc1000289 to 1000292. LAOS • ♀; “ Xieng Khouang, 8.i.1919; R.V. de Salvaza; Indo-China, B.M. 1924-315; BMNH (E) 1237162, Southeast Asia”. MYANMAR • ♂; “ex Mus. Calcutta, B.M. 1930–1. Man Hpat, N. Tawngpeng, N. Shan States, Upper Burma, 11-ii-15., 4600 ft.; J. Coggin Brown; in dung; Southeast Asia;” BMNH (E) 1237161 • ♀; “ Loi, Tawng Kyan, Tawng, N. Shan States, Upper Burma • ♂; Tawng Peng, Pansai, N.Shan States, Upper Burma, 4660 ft. ”. Description MEASUREMENTS (in mm). TL=15–20.5; BW= 8.5–11.5. Male Black, subopaque; head, lower surface and legs dark red; pygidium lower surface and fringe of elytra with reddish hairs; body oval, compact, convex; clypeus elliptical, finely, closely rugulose; forehead rugosely punctured with sharply elevated carina; clypeal carina short, transverse, slightly curved; vertex with a short transverse curved carina before carinate posterior margin of head; pronotum strongly, closely punctured, with feeble median groove, base margined, basal foveae very deep, space between them finely rugose; elytra very convex, opaque, finely striate with intervals carinate along middle; pygidium feebly rugulose, thinly clothed with erect, long reddish hairs; metasternum granular, broadly grooved in the middle; abdomen hairy at sides, smooth in the middle; profemur with sharp tooth near middle of front edge; protibia with finely serrate carina beneath and a tooth near base; middle coxa toothed at its posterior end. AEDEAGUS (in mm) (Fig. 11E–H). LP=2.86, Lp =2.68; BP= 1.25, BpB=1.07, BpT=0.36. Phallobase and parameres almost of same size; parameres straight above and slightly curved below with the tips blunt and rounded. Female Clypeus extended and reflexed anteriorly, slightly notched; no sharp tooth on underside of protibia and profemur. Distribution India: Assam, Arunachal Pradesh, Meghalaya, Mizoram; Other countries: China; Laos; Myanmar; Thailand (Singh et al. 2010; Gupta et al. 2015; Bezděk 2016; Schoolmeesters 2022).

Published as part of Karimbumkara, Seena Narayanan & Priyadarsanan, Dharma Rajan, 2024, Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India, pp. 1-61 in European Journal of Taxonomy 956 on pages 23-24, DOI: 10.5852/ejt.2024.956.2657, http://zenodo.org/record/13742850