Fig. 4 Wall-structure of Zagrosella rigaudii n.gen., n.sp. from the upper Maastrichtian of SW Iran (a–b) and Biokovina gradacensis Gušić from the Liassic of Croatia (c–d). Polygonal parapores are marked with an arrow in a. Thin-sections: 2NG 27 (a), 2NG 87 (b). Remarks and comparisons: The wall structure of Zag- the Late Cretaceous, it can be compared to some extent rosella with its "uniform, parallel, radial elements cov- (e.g., similar wall-structure, strengthenings) with Braciered by some kind of tectum" (Hottinger, 2006, p. 29) can ana Schlagintweit & Cvetko-Tesovic, 2016 (Santonian?- be compared with the keriotheca-like (or pseudokerithe- lower Campanian of Croatia). The latter differs from cal) texture of some Mesozoic larger benthic foraminifera Zagrosella n.gen. above all by its test morphology (elon- (e.g., Schroeder et al., 1975; De Castro, 1981; Septfon- gate compressed), and the foraminal characteristics (sevtaine, 1981; Banner et al., 1991; Rigaud et al., 2015). eral close-set openings, equal in diameter to the wall al- Regarding differences to parapores (or canaliculi), the veolae, and may have short peristomal rims at their mardifferences are vague and not clearly delimited (Hot- gins) (see Schlagintweit and Cvetko-Tešović, 2016 for tinger, 2006: "usually much larger"…"often more irregu- details). lar"). Zagrosella n.gen. is morphologically very similar to Last but not least we note morphological similarities the Liassic Biokovina Gušić, 1977, both monospecific to the Late Jurassic Labyrinthina Weynschenk that differs genera. The wall structure of Zagrosella appears more above all from Zagrosiella by its wall being "agglutinatevolved with larger parapores (diameter 0.01 to 0.03 mm, ed, simple in structure, microgranular, imperforate" see description below) whereas in Biokovina these are (Loeblich and Tappan (1987, p. 96). thinner (but with overlapping range), and bifurcating (diameter ~0.01 to 0.15 mm). There are also differences in Zagrosella rigaudii Schlagintweit & Rashidi, n.sp. the endoskeletal structures, being more massive (in the Figs. 3 pars, 4a–b, 5–7 central part of the test) and may fuse in Biokovina where- Origin of the name: The species name refers to Sylvain as in Zagrosella they occur preferentially on the sides of Rigaud (Singapore) for his outstanding contributions to the chambers, sometimes only partially differentiated the micropalaeontology and phylogeny of benthic foramfrom the chamber wall, and never fused. They are here inifera. interpreted as pillars (originating from the septa), not as Holotype: Slightly oblique equatorial section illustrated septula (originating from the wall) (see Hottinger, 2006) in Figure 5A, thin-section NG 88. or strengthenings (see Rigaud et al., 2013). Another mor- Paratypes: Figs. 5B–G, 6, 7A–B. phologically similar genus of the family Biokovinidae is Description: Test free, lenticular, with oscillating plane Bosniella Gušić, 1977. It differs from Zagrosella above of coiling in the early stage, later planispiral, and finally all by its chambers lacking any endoskeletal structures. In maybe uncoiling, rectilinear. It is compressed axially,

Published as part of Schlagintweit, Felix & Rashidi, Koorosh, 2017, Zagrosella Rigaudii N. Gen., N. Sp., A New Biokovinoidean Foraminifer From The Maastrichtian Of Iran, pp. 3-13 in Acta Palaeontologica Romaniae 13 (1) on page 7, DOI: 10.5281/zenodo.13189884