1. Macrozamia Moorei bears numerous lateral cones in the axils of leaves, in this respect being identical with the mesozoic Bennettitales. 2. The pollen grain, at the time of shedding, contains a persistent prothallial cell, a generative cell, and a tube cell; the exine does not cover the apical part of the grain. In the young pollen tube the generative cell has given rise to a stalk cell and a body cell like those of other cycads. 3. The archegonial chamber is unusually deep, and the archegonia are of moderate size. 4. The embryogeny resembles that of Cycas and differs from that of Zamia and Ceratozamia. 5. The species, which represents the nearest approach to the mesozoic Bennettitales, is in immediate danger of extinction.