Clusia and Clusiodes Clusia. Third stage larva: length 8–9mm; maximum width 1.5mm; maximum height 1.6mm; puparium: length 4.5–8mm and pupal spiracles present; anterior spiracles with 4–6 respiratory bulbs (Figs 8–9); antero-dorsal margins of metathorax (Fig 2) and abdominal segments 1–6 with a narrow band of spiculate ridges (Fig 20) extending transversely and ending before the lateral margins, bands becoming narrower and spicule rows less frequent from segment 1 to 6; locomotory organs on the anterior margin of the metathorax and abdominal segments 1–7 widest medially, tapering laterally with even-sized spicules (Fig 22); margins of the anal plate with numerous spiculate ridges (Fig 56); middle region of the anal segment with a complete, large and conspicuous flange Figs 26–27); hooks associated with posterior spiracles with incurved apices (Fig 24); posterior spiracular plates elevated and pale in the larva, black in the puparium with extensive interspiracular setae bearing 5–15 branches (often broken or missing) (Fig 24). Clusiodes. Third stage larva: length 6–8mm; maximum width 0.8mm; maximum height 1mm; puparium: length 3.5–5mm, pupal spiracles may or may not be present, if present often expanded imperfectly; anterior spiracles with 3–10 respiratory bulbs (Figs 10–18); antero-dorsal margins of metathorax (Fig 1) and abdominal segments 1–2 with a narrow band of smooth ridges lacking spicules (Fig 19), extending transversely and ending before the lateral margins; locomotory organs on the anterior margin of the metathorax and abdominal segments 1–7 widest medially, tapering laterally with variable-sized spicules: anterior band with rows of ridges bearing small spicules; next, a narrow band of ridges with large spicules followed by a wider band of smooth, non-spiculate ridges (Fig 21); margins of the anal plate with few spiculate ridges (Fig 57); middle region of the anal segment lacking complete circumventing flanges, usually several, incomplete and small folds or wrinkles present or wrinkles more or less absent; apices of hooks associated with posterior spiracles orientated dorsally (Fig 23); posterior spiracular plates pale in the larva and puparium, not elevated and interspiracular setae with up to 5 terminal branches (often broken or missing) (Fig 23).

Published as part of Horsfield, David, 2013, Development sites and early stages of eleven species of Clusiidae (Diptera) occurring in Europe, pp. 401-427 in Zootaxa 3619 (4) on page 413, DOI: 10.11646/zootaxa.3619.4.1, http://zenodo.org/record/223688