Available information on seed dormancy for various members of the Saxifragales and phylogenetic relationships within this order allowed us to accurately predict that Daphniphyllum glaucescens seeds have morphophysiological dormancy (MPD). However, our hypothesis that seeds had deep simple epicotyl MPD, i.e., nondeep physiological dormancy (PD) in root and deep PD in shoot, was not supported. Both the root and the shoot (cotyledons) of the underdeveloped embryo of D. glaucescens have nondeep PD. Exposure to moderate (15°/6°, 20°/10°C), rather than high (25°/15°C), temperatures for 10–12 wk broke the PD of the hypocotyl/root. After hypocotyl emergence, seeds with an attached developing root system did not require cold stratification to break the PD of the shoot. The PD of the shoot was broken by an additional 10–12 wk at moderate temperatures, during which time cotyledons slowly grew inside the seed. As cotyledons grew, all of the hypocotyl was pushed out of the seed, and the final cotyledon length was almo...