Abstract The paper records the existence of Angiosperins in England in Aptian times, i.e. at a geological period when they have been hitherto supposed not to exist in Northern Europe; describes botanically the anatomy of these fossils, which come under three new genera; and notes the points of structural and phylogenetic interest in them, as being the oldest Angiosperms of which the anatomy is preserved, and contemporaneous with Bennettites.